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Rolle's rock-cress

littleleaf suncress, Macoun's suncress, sloppy suncress

Habit Perennials; long-lived; sexual; caudex woody. Perennials; usually long-lived; apomictic; caudex often woody.
Stems

usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 1.5–2.5 dm, glabrous throughout.

1 or several per caudex branch, arising from center or margin of rosette near ground surface, 2–3.6 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, 0.1–0.8 mm, glabrous distally.

Basal leaves

blade oblanceolate, 3–8 mm wide, margins entire, ciliate proximally, trichomes (simple), 0.2–0.7 mm, surfaces glabrous or sparsely pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.4 mm.

blade oblanceolate to linear-oblanceolate, 1–3(–5) mm wide, margins entire or minutely toothed, ciliate proximally, trichomes (simple), to 0.7 mm, surfaces densely pubescent, trichomes short-stalked, 3–6-rayed, 0.07–0.3 mm.

Cauline leaves

6–12, sometimes concealing stem proximally;

blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous.

(5–)9–16, often concealing stem proximally;

blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous.

Racemes

3–7-flowered, unbranched.

10–33-flowered, usually unbranched.

Flowers

divaricate-ascending at anthesis;

sepals glabrous;

petals creamy white, 8–11 × 2–2.5 mm, glabrous;

pollen ellipsoid.

divaricate-ascending at anthesis;

sepals pubescent;

petals lavender to purple, 4–5 × 0.5–0.8 mm, glabrous;

pollen spheroid.

Fruiting pedicels

strongly curved proximally, 4–6 mm, glabrous.

divaricate-ascending to horizontal, straight, 5–13(–17) mm, usually densely pubescent, rarely glabrous, trichomes appressed, 2- or 3-rayed.

Fruits

pendent, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), 3.5–6 cm × 2.5–3.5 mm;

valves glabrous;

ovules 14–22 per ovary;

style 0.5–1 mm.

divaricate-ascending to descending, not appressed to rachis, not secund, usually curved, edges parallel, 3.5–6.5 cm × 0.8–1.2 mm;

valves glabrous;

ovules 56–78 per ovary;

style 0.1–0.4 mm.

Seeds

uniseriate, 3–4 × 1.5–2 mm;

wing distal and proximal, 0.3–0.6 mm wide.

uniseriate, 1–1.2 × 0.7–1 mm;

wing often continuous, 0.05–0.15 mm wide.

Boechera rollei

Boechera macounii

Phenology Flowering Aug. Flowering May–Jul.
Habitat Peridotite rocks on sparsely forested slopes Rocky hillsides in open pinewoods
Elevation 1600-1800 m (5200-5900 ft) 2200-2900 m (7200-9500 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
ID; MT; WY; BC
Discussion

Boechera rollei is easily distinguished from the habitally similar B. suffrutescens by its longer (8–11 versus 4.5–6 mm) petals, narrower (2.5–3.5 versus 3–6 mm) fruits, and narrower (0.3–0.6 versus 0.8–1.5 mm) seed wings. This distinctive sexual species is known only from the vicinity of the type locality on the divide between the Applegate and Klamath Rivers in Siskiyou County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera macounii is an apomictic species that arose through hybridization between B. collinsii and B. microphylla (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 403. FNA vol. 7, p. 389.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis rollei Arabis macounii, Arabis densicaulis, Arabis microphylla var. macounii
Name authority (Rollins) Al-Shehbaz: Novon 13: 389. (2003) (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 267. (2007)
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