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Rolle's rock-cress

Habit Perennials; long-lived; sexual; caudex woody. Biennials or perennials; short-lived; apomictic or sexual; caudex present or absent.
Stems

usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 1.5–2.5 dm, glabrous throughout.

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6.7 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade oblanceolate, 3–8 mm wide, margins entire, ciliate proximally, trichomes (simple), 0.2–0.7 mm, surfaces glabrous or sparsely pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.4 mm.

blade oblanceolate, 2–5 mm wide, margins entire, sometimes ciliate near petiole base, trichomes (usually simple), to 0.7 mm, surfaces sparsely to moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm.

Cauline leaves

6–12, sometimes concealing stem proximally;

blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous.

7–40, often concealing stem proximally;

blade auricles 1–4 mm, surfaces of distalmost leaves glabrous or sparsely pubescent.

Racemes

3–7-flowered, unbranched.

10–60-flowered, usually unbranched.

Flowers

divaricate-ascending at anthesis;

sepals glabrous;

petals creamy white, 8–11 × 2–2.5 mm, glabrous;

pollen ellipsoid.

divaricate to pendent at anthesis;

sepals pubescent;

petals white, 4–8 × 1–2 mm, glabrous;

pollen ellipsoid or spheroid.

Fruiting pedicels

strongly curved proximally, 4–6 mm, glabrous.

descending to reflexed, most abruptly recurved near base, 5–10 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

Fruits

pendent, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), 3.5–6 cm × 2.5–3.5 mm;

valves glabrous;

ovules 14–22 per ovary;

style 0.5–1 mm.

descending to reflexed, not appressed to rachis, strongly secund, often curved, edges parallel, 0.35–0.65 cm × (2–)2.2–2.5 mm;

valves glabrous;

ovules 80–102 per ovary.

Seeds

uniseriate, 3–4 × 1.5–2 mm;

wing distal and proximal, 0.3–0.6 mm wide.

uniseriate to sub-biseriate, 1.4–1.7 × 1–1.2 mm;

wing continuous, to 0.2 mm wide.

2n

= 14, 21.

Boechera rollei

Boechera holboellii

Phenology Flowering Aug. Flowering Jul–Aug.
Habitat Peridotite rocks on sparsely forested slopes Rocky slopes and gravelly soil near coast
Elevation 1600-1800 m (5200-5900 ft) 100-300 m (300-1000 ft)
Distribution
from FNA
CA
[BONAP county map]
from USDA
Greenland
[BONAP county map]
Discussion

Boechera rollei is easily distinguished from the habitally similar B. suffrutescens by its longer (8–11 versus 4.5–6 mm) petals, narrower (2.5–3.5 versus 3–6 mm) fruits, and narrower (0.3–0.6 versus 0.8–1.5 mm) seed wings. This distinctive sexual species is known only from the vicinity of the type locality on the divide between the Applegate and Klamath Rivers in Siskiyou County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Both sexual diploid and apomictic triploid collections of Boechera holboellii are known; available data suggest that the triploid is an autopolyploid derivative of the sexual diploid but further study is needed. In the narrow circumscription adopted here, it appears to be restricted to Greenland (see discussion of B. quebecensis in M. D. Windham and I. A. Al-Shehbaz 2007b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 403. FNA vol. 7, p. 384.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis rollei Arabis holboellii, Arabis holboellii var. tenuis, B. tenuis
Name authority (Rollins) Al-Shehbaz: Novon 13: 389. (2003) (Hornemann) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976)
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