Boechera rollei |
Boechera harrisonii |
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Rolle's rock-cress |
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Habit | Perennials; long-lived; sexual; caudex woody. | Perennials; long-lived; (some-what cespitose); apomictic; caudex often woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 1.5–2.5 dm, glabrous throughout. |
usually 3–7 per caudex branch, arising from margin of rosette near ground surface, 0.5–2.5 dm, usually sparsely pubescent proximally, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.06–0.2 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 3–8 mm wide, margins entire, ciliate proximally, trichomes (simple), 0.2–0.7 mm, surfaces glabrous or sparsely pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.4 mm. |
blade narrowly oblanceolate, 2–4 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 0.5 mm, surfaces sparsely pubescent, trichomes short-stalked, 2–5-rayed, 0.1–0.25 mm. |
Cauline leaves | 6–12, sometimes concealing stem proximally; blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous. |
3–7, not concealing stem; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 3–7-flowered, unbranched. |
5–12-flowered, usually unbranched. |
Flowers | divaricate-ascending at anthesis; sepals glabrous; petals creamy white, 8–11 × 2–2.5 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender, 5–7.5 × 0.8–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | strongly curved proximally, 4–6 mm, glabrous. |
divaricate-ascending, straight, 8–12 mm, glabrous. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), 3.5–6 cm × 2.5–3.5 mm; valves glabrous; ovules 14–22 per ovary; style 0.5–1 mm. |
usually divaricate-ascending, rarely nearly horizontal, not appressed to rachis, not secund, curved to straight, edges parallel, 3–4.7 cm × 1–1.5 mm; valves glabrous; ovules 64–86 per ovary; style 0.2–0.5 mm. |
Seeds | uniseriate, 3–4 × 1.5–2 mm; wing distal and proximal, 0.3–0.6 mm wide. |
uniseriate, 1.1–1.3 × 0.8–1.1 mm; wing continuous, 0.08–0.1 mm wide. |
Boechera rollei |
Boechera harrisonii |
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Phenology | Flowering Aug. | Flowering May–Jun. |
Habitat | Peridotite rocks on sparsely forested slopes | Limestone and quartzite cliffs |
Elevation | 1600-1800 m (5200-5900 ft) | 1500-1600 m (4900-5200 ft) |
Distribution |
CA |
UT |
Discussion | Boechera rollei is easily distinguished from the habitally similar B. suffrutescens by its longer (8–11 versus 4.5–6 mm) petals, narrower (2.5–3.5 versus 3–6 mm) fruits, and narrower (0.3–0.6 versus 0.8–1.5 mm) seed wings. This distinctive sexual species is known only from the vicinity of the type locality on the divide between the Applegate and Klamath Rivers in Siskiyou County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera harrisonii is an apomictic species that arose through hybridization between B. microphylla and B. perennans (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). Boechera harrisonii is known only from Utah and Wasatch counties in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 403. | FNA vol. 7, p. 383. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis rollei | Arabis harrisonii, B. microphylla var. harrisonii |
Name authority | (Rollins) Al-Shehbaz: Novon 13: 389. (2003) | (S. L. Welsh) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) |
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