Boechera repanda |
Boechera williamsii |
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Greene's rockcress, Yosemite rockcress |
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Habit | Perennials; short- to long-lived; sexual; caudex usually not woody. | Perennials; short-lived; sexual; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–9 dm, densely pubescent proximally, trichomes short- to long-stalked, 2–6-rayed, 0.2–0.5 mm, rarely mixed with simple ones, to 1.5 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
Basal leaves | blade broadly oblanceolate to obovate, 7–25(–50) mm wide, margins usually repand to coarsely dentate, rarely entire, not ciliate, surfaces moderately to densely pubescent, trichomes short- to long-stalked, 3–6-rayed, 0.2–0.5 mm. |
blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
Cauline leaves | (3–)8–30, not concealing stem; blade auricles absent, surfaces of distalmost leaves sparsely pubescent. |
6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 7–25-flowered, sometimes branched. |
5–17-flowered, unbranched. |
Flowers | erect to ascending at anthesis; sepals glabrous or sparsely pubescent; petals white, 3.5–6 × 0.8–1 mm, glabrous; pollen ellipsoid. |
erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | erect to divaricate-ascending, straight, 3–10 mm, glabrous or pubescent, trichomes spreading, usually 2–5-rayed, rarely simple. |
erect, straight, 2–4(–7) mm, glabrous. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (3.5–)5–13.5 cm × 2.5–4 mm; valves glabrous or pubescent throughout; ovules 34–50 per ovary; style 0.5–1.5 mm. |
erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 2.5–5(–6) × 2–3(–3.5) mm; wing continuous, 0.7–1.8 mm wide. |
sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
2n | = 14. |
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Boechera repanda |
Boechera williamsii |
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Phenology | Flowering Jun–Jul. | Flowering May–Jun. |
Habitat | Rock outcrops, talus, and gravelly soil in meadows and open pine forests | Gravelly soil in sagebrush-grassland communities |
Elevation | 1600-3500 m (5200-11500 ft) | 2300-2800 m (7500-9200 ft) |
Distribution |
CA; NV
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WY |
Discussion | Boechera repanda is a sexual diploid that is distinguished from all other western North American species by its wide (10–50 mm), repand to coarsely dentate basal leaves. The form recognized as var. greenei appears to be a growth response to ecologically marginal habitats. Boechera repanda occurs sporadically in the Sierra Nevada and Transverse Ranges of California, and barely extends into the Carson Range in western Nevada. Arabis inamoena Greene (1911), not Greene (1908) is an illegitimate name, sometimes found in synonymy with Boechera repanda. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 402. | FNA vol. 7, p. 411. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis repanda, Arabis repanda var. greenei | Arabis williamsii |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 388. (2003) | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) |
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