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Greene's rockcress, Yosemite rockcress

spoonleaf rockcress

Habit Perennials; short- to long-lived; sexual; caudex usually not woody. Perennials; short- to long-lived; sexual; caudex usually not woody (rarely with persistent, crowded leaf bases).
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–9 dm, densely pubescent proximally, trichomes short- to long-stalked, 2–6-rayed, 0.2–0.5 mm, rarely mixed with simple ones, to 1.5 mm, glabrous or sparsely pubescent distally.

simple or few to several per caudex branch, arising from center of rosette near ground surface, 1.5–3.5(–5) dm, densely pubescent proximally, trichomes simple mixed with short-stalked ones, 2-rayed, 0.3–0.7 mm.

Basal leaves

blade broadly oblanceolate to obovate, 7–25(–50) mm wide, margins usually repand to coarsely dentate, rarely entire, not ciliate, surfaces moderately to densely pubescent, trichomes short- to long-stalked, 3–6-rayed, 0.2–0.5 mm.

blade narrowly oblanceolate, 1.5–3(–4) mm wide, margins entire, strongly ciliate at least along petiole, trichomes (simple), to 1 mm, surfaces glabrous or sparsely pubescent, trichomes simple and short-stalked, 2-rayed, 0.3–0.7 mm.

Cauline leaves

(3–)8–30, not concealing stem;

blade auricles absent, surfaces of distalmost leaves sparsely pubescent.

5–15(–20), often concealing stem proximally;

blade auricles 0.5–1.5 mm, surfaces of distalmost leaves usually glabrous.

Racemes

7–25-flowered, sometimes branched.

10–30-flowered, usually unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous or sparsely pubescent;

petals white, 3.5–6 × 0.8–1 mm, glabrous;

pollen ellipsoid.

divaricate-ascending at anthesis;

sepals pubescent;

petals usually white, rarely pale lavender, 3–3.7(–4) × 0.5–0.8 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

erect to divaricate-ascending, straight, 3–10 mm, glabrous or pubescent, trichomes spreading, usually 2–5-rayed, rarely simple.

horizontal or slightly descending, curved or angled downward, 6–10(–15) mm, usually glabrous, rarely sparsely pubescent, trichomes spreading, simple.

Fruits

divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (3.5–)5–13.5 cm × 2.5–4 mm;

valves glabrous or pubescent throughout;

ovules 34–50 per ovary;

style 0.5–1.5 mm.

pendent, not appressed to rachis, rarely slightly secund, straight or gently curved, edges parallel, 3.3–5.7 cm × 1.2–1.8 mm;

valves glabrous;

ovules 90–126 per ovary;

style 0.1–0.4 mm.

Seeds

uniseriate, 2.5–5(–6) × 2–3(–3.5) mm;

wing continuous, 0.7–1.8 mm wide.

biseriate, 0.7–0.9 × 0.5–0.6 mm;

wing distal or, sometimes, absent, 0.05–0.1 mm wide.

2n

= 14.

= 14.

Boechera repanda

Boechera spatifolia

Phenology Flowering Jun–Jul. Flowering May–Aug.
Habitat Rock outcrops, talus, and gravelly soil in meadows and open pine forests Rocky slopes and gravelly soil in sagebrush, pinyon-juniper woodlands, open conifer forests and subalpine meadows
Elevation 1600-3500 m (5200-11500 ft) 1800-2800 m (5900-9200 ft)
Distribution
from FNA
CA; NV
[WildflowerSearch map]
from FNA
CO; NM
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera repanda is a sexual diploid that is distinguished from all other western North American species by its wide (10–50 mm), repand to coarsely dentate basal leaves. The form recognized as var. greenei appears to be a growth response to ecologically marginal habitats. Boechera repanda occurs sporadically in the Sierra Nevada and Transverse Ranges of California, and barely extends into the Carson Range in western Nevada.

Arabis inamoena Greene (1911), not Greene (1908) is an illegitimate name, sometimes found in synonymy with Boechera repanda.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera spatifolia is a sexual diploid that usually has been treated as a variety of Arabis (Boechera) fendleri but appears to be sufficiently distinct to warrant recognition at species level (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). There is little geographic overlap between the two, with B. spatifolia confined to the mountains of central Colorado and north-central New Mexico and B. fendleri ranging from western New Mexico and the Four Corners region through northern Arizona to southern Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 402. FNA vol. 7, p. 408.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis repanda, Arabis repanda var. greenei Arabis spatifolia, Arabis fendleri var. spatifolia, B. fendleri subsp. spatifolia, B. fendleri var. spatifolia
Name authority (S. Watson) Al-Shehbaz: Novon 13: 388. (2003) (Rydberg) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 84. (2006)
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