Boechera repanda |
Boechera pauciflora |
|
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Greene's rockcress, Yosemite rockcress |
Columbia rockcress, few-flower rock-cress, fuzzy suncress, hairy-stem rock-cress, small-flower rockcress |
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Habit | Perennials; short- to long-lived; sexual; caudex usually not woody. | Perennials; short- to long-lived; apomictic; caudex sometimes woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–9 dm, densely pubescent proximally, trichomes short- to long-stalked, 2–6-rayed, 0.2–0.5 mm, rarely mixed with simple ones, to 1.5 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.4–)3–11.2 dm, densely pubescent proximally, trichomes simple, 0.6–1.5 mm, mixed with stalked, 2- (or 3-)rayed ones, 0.2–0.4 mm, sparsely pubescent distally. |
Basal leaves | blade broadly oblanceolate to obovate, 7–25(–50) mm wide, margins usually repand to coarsely dentate, rarely entire, not ciliate, surfaces moderately to densely pubescent, trichomes short- to long-stalked, 3–6-rayed, 0.2–0.5 mm. |
blade oblanceolate, 3–10 mm wide, margins usually dentate, rarely entire, sometimes ciliate, trichomes (simple or branched), to 1 mm, surfaces densely pubescent, trichomes stalked, 2–5-rayed, 0.3–0.6 mm. |
Cauline leaves | (3–)8–30, not concealing stem; blade auricles absent, surfaces of distalmost leaves sparsely pubescent. |
(8–)14–60, often concealing stem proximally; blade auricles (1–)3–10 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 7–25-flowered, sometimes branched. |
17–60-flowered, usually unbranched. |
Flowers | erect to ascending at anthesis; sepals glabrous or sparsely pubescent; petals white, 3.5–6 × 0.8–1 mm, glabrous; pollen ellipsoid. |
divaricate at anthesis; sepals pubescent; petals lavender to whitish, 5–8 × 1–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect to divaricate-ascending, straight, 3–10 mm, glabrous or pubescent, trichomes spreading, usually 2–5-rayed, rarely simple. |
horizontal to divaricate-descending, usually straight, rarely slightly recurved, 4–13 mm, pubescent, trichomes spreading, 2- or 3-rayed. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (3.5–)5–13.5 cm × 2.5–4 mm; valves glabrous or pubescent throughout; ovules 34–50 per ovary; style 0.5–1.5 mm. |
horizontal, divaricate-descending or widely pendent, not appressed to rachis, not secund, curved, edges parallel, 5.5–10.5 cm × 1.5–2.2 mm; valves glabrous; ovules 80–162 per ovary; style 0.05–0.5 mm. |
Seeds | uniseriate, 2.5–5(–6) × 2–3(–3.5) mm; wing continuous, 0.7–1.8 mm wide. |
uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.25 mm wide. |
2n | = 14. |
= 21. |
Boechera repanda |
Boechera pauciflora |
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Phenology | Flowering Jun–Jul. | Flowering May–Jun. |
Habitat | Rock outcrops, talus, and gravelly soil in meadows and open pine forests | Rocky soil in sagebrush areas, mountain shrub communities, edges of conifer forests |
Elevation | 1600-3500 m (5200-11500 ft) | 600-2500 m (2000-8200 ft) |
Distribution |
CA; NV
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CA; ID; MT; NV; OR; UT; WA; WY; BC
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Discussion | Boechera repanda is a sexual diploid that is distinguished from all other western North American species by its wide (10–50 mm), repand to coarsely dentate basal leaves. The form recognized as var. greenei appears to be a growth response to ecologically marginal habitats. Boechera repanda occurs sporadically in the Sierra Nevada and Transverse Ranges of California, and barely extends into the Carson Range in western Nevada. Arabis inamoena Greene (1911), not Greene (1908) is an illegitimate name, sometimes found in synonymy with Boechera repanda. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera pauciflora is an apomictic species that arose through hybridization between B. retrofracta and B. sparsiflora. Specimens of B. pauciflora are commonly identified as Arabis holboellii var. pinetorum (= B. pinetorum), a superficially similar species restricted to the northern Sierra Nevada and southern Cascade Range (see M. D. Windham and I. A. 2007 for detailed comparison). Arabis elegans A. Nelson (1900), not Tineo & Lojacono (1886) is an illegitimate name, sometimes found in synonymy with Boechera pauciflora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 402. | FNA vol. 7, p. 393. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis repanda, Arabis repanda var. greenei | Sisymbrium pauciflorum, Arabis arcuata var. subvillosa, Arabis columbiana, Arabis perelegans, Arabis sparsiflora var. columbiana, Arabis sparsiflora var. subvillosa, B. sparsiflora var. subvillosa |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 388. (2003) | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 268. (2007) |
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