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Greene's rockcress, Yosemite rockcress

Crater Lake rock-cress

Habit Perennials; short- to long-lived; sexual; caudex usually not woody. Perennials; long-lived; (cespitose); apomictic; caudex woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–9 dm, densely pubescent proximally, trichomes short- to long-stalked, 2–6-rayed, 0.2–0.5 mm, rarely mixed with simple ones, to 1.5 mm, glabrous or sparsely pubescent distally.

usually 1 per caudex branch, arising from center of rosette, near ground surface or somewhat elevated on woody base, or 1–3.5 dm, sparsely pubescent proximally, trichomes short-stalked, 3–6-rayed, 0.1–0.3 mm, glabrous distally.

Basal leaves

blade broadly oblanceolate to obovate, 7–25(–50) mm wide, margins usually repand to coarsely dentate, rarely entire, not ciliate, surfaces moderately to densely pubescent, trichomes short- to long-stalked, 3–6-rayed, 0.2–0.5 mm.

blade narrowly oblanceolate, 1–5 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 3–6-rayed, 0.1–0.3 mm.

Cauline leaves

(3–)8–30, not concealing stem;

blade auricles absent, surfaces of distalmost leaves sparsely pubescent.

3–13, often concealing stem proximally;

blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous.

Racemes

7–25-flowered, sometimes branched.

5–32-flowered, usually unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous or sparsely pubescent;

petals white, 3.5–6 × 0.8–1 mm, glabrous;

pollen ellipsoid.

divaricate-ascending at anthesis;

sepals pubescent;

petals lavender to purple, 5–6 × 1.5–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

erect to divaricate-ascending, straight, 3–10 mm, glabrous or pubescent, trichomes spreading, usually 2–5-rayed, rarely simple.

horizontal to descending, straight or slightly curved downward, 4–11 mm, glabrous.

Fruits

divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (3.5–)5–13.5 cm × 2.5–4 mm;

valves glabrous or pubescent throughout;

ovules 34–50 per ovary;

style 0.5–1.5 mm.

horizontal or descending, not appressed to rachis, secund, straight, edges slightly undulate (not parallel), 2–4 cm × 2–3 mm;

valves glabrous;

ovules 40–54 per ovary;

style 0.2–0.5 mm.

Seeds

uniseriate, 2.5–5(–6) × 2–3(–3.5) mm;

wing continuous, 0.7–1.8 mm wide.

uniseriate, 2–2.5 × 1.7–2 mm;

wing continuous, 0.5–1 mm wide.

2n

= 14.

Boechera repanda

Boechera horizontalis

Phenology Flowering Jun–Jul. Flowering Jul–Aug.
Habitat Rock outcrops, talus, and gravelly soil in meadows and open pine forests Dry pumice slopes
Elevation 1600-3500 m (5200-11500 ft)
Distribution
from FNA
CA; NV
[WildflowerSearch map]
from FNA
OR
[BONAP county map]
Discussion

Boechera repanda is a sexual diploid that is distinguished from all other western North American species by its wide (10–50 mm), repand to coarsely dentate basal leaves. The form recognized as var. greenei appears to be a growth response to ecologically marginal habitats. Boechera repanda occurs sporadically in the Sierra Nevada and Transverse Ranges of California, and barely extends into the Carson Range in western Nevada.

Arabis inamoena Greene (1911), not Greene (1908) is an illegitimate name, sometimes found in synonymy with Boechera repanda.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera horizontalis is an apomictic species that arose through hybridization between B. lemmonii and B. suffrutescens (M. D. Windham and I. A. Al-Shehbaz 2007). Boechera horizontalis is known only from the vicinity of Crater Lake in south-central Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 402. FNA vol. 7, p. 384.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis repanda, Arabis repanda var. greenei Arabis horizontalis, Arabis suffrutescens var. horizontalis
Name authority (S. Watson) Al-Shehbaz: Novon 13: 388. (2003) (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007)
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