Boechera repanda |
Boechera fructicosa |
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Greene's rockcress, Yosemite rockcress |
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Habit | Perennials; short- to long-lived; sexual; caudex usually not woody. | Perennials; short-lived; apomictic; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–9 dm, densely pubescent proximally, trichomes short- to long-stalked, 2–6-rayed, 0.2–0.5 mm, rarely mixed with simple ones, to 1.5 mm, glabrous or sparsely pubescent distally. |
3 to many per caudex branch, arising from margin of rosette near ground surface, 3.5–6 dm, pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.2–0.3 mm, glabrous distally. |
Basal leaves | blade broadly oblanceolate to obovate, 7–25(–50) mm wide, margins usually repand to coarsely dentate, rarely entire, not ciliate, surfaces moderately to densely pubescent, trichomes short- to long-stalked, 3–6-rayed, 0.2–0.5 mm. |
blade oblanceolate, 3–7 mm wide, margins dentate, ciliate proximally, trichomes (simple), to 0.3 mm, surfaces pubescent, trichomes short-stalked, 4–8-rayed, 0.05–0.2 mm. |
Cauline leaves | (3–)8–30, not concealing stem; blade auricles absent, surfaces of distalmost leaves sparsely pubescent. |
5–10, not concealing stem; blade auricles 0.5–1 mm, surfaces of distalmost leaves glabrous. |
Racemes | 7–25-flowered, sometimes branched. |
10–18-flowered, usually unbranched. |
Flowers | erect to ascending at anthesis; sepals glabrous or sparsely pubescent; petals white, 3.5–6 × 0.8–1 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals lavender, 4–6 × 1–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect to divaricate-ascending, straight, 3–10 mm, glabrous or pubescent, trichomes spreading, usually 2–5-rayed, rarely simple. |
divaricate-descending, straight, 5–10 mm, glabrous. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (3.5–)5–13.5 cm × 2.5–4 mm; valves glabrous or pubescent throughout; ovules 34–50 per ovary; style 0.5–1.5 mm. |
divaricate-descending, not appressed to rachis, not secund, curved, edges parallel, 4–6 cm × 1.5–1.8 mm; valves glabrous; ovules 60–70 per ovary; style 0.1–0.4 mm. |
Seeds | uniseriate, 2.5–5(–6) × 2–3(–3.5) mm; wing continuous, 0.7–1.8 mm wide. |
uniseriate, 1.5–2 × 1–1.3 mm; wing distal, ca. 0.1 mm wide. |
2n | = 14. |
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Boechera repanda |
Boechera fructicosa |
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Phenology | Flowering Jun–Jul. | Flowering Jun–Jul. |
Habitat | Rock outcrops, talus, and gravelly soil in meadows and open pine forests | Dry, disturbed soil |
Elevation | 1600-3500 m (5200-11500 ft) | ca. 2000 m (ca. 6600 ft) |
Distribution |
CA; NV
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WY |
Discussion | Boechera repanda is a sexual diploid that is distinguished from all other western North American species by its wide (10–50 mm), repand to coarsely dentate basal leaves. The form recognized as var. greenei appears to be a growth response to ecologically marginal habitats. Boechera repanda occurs sporadically in the Sierra Nevada and Transverse Ranges of California, and barely extends into the Carson Range in western Nevada. Arabis inamoena Greene (1911), not Greene (1908) is an illegitimate name, sometimes found in synonymy with Boechera repanda. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera fructicosa is known only from the type collection (A. & E. Nelson 5681) taken near Undine Falls in Park County. Multiple plants are represented (comprising the holotype and four isotypes), though one specimen with the same number at RM is B. stricta. Although the exact parentage of this apomictic hybrid remains in doubt, it is virtually certain that B. microphylla contributed at least one genome. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 402. | FNA vol. 7, p. 379. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis repanda, Arabis repanda var. greenei | Arabis fructicosa |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 388. (2003) | (A. Nelson) Al-Shehbaz: Novon 13: 385. (2003) |
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