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Greene's rockcress, Yosemite rockcress

Brewer's rock-cress

Habit Perennials; short- to long-lived; sexual; caudex usually not woody. Perennials; long-lived; sexual; caudex woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–9 dm, densely pubescent proximally, trichomes short- to long-stalked, 2–6-rayed, 0.2–0.5 mm, rarely mixed with simple ones, to 1.5 mm, glabrous or sparsely pubescent distally.

usually 1 per caudex branch, arising from center of rosette, somewhat elevated on woody base or from ground surface, 0.6–3.5(–4.5) dm, densely pubescent proximally, trichomes long-stalked and simple, 2-rayed (sometimes all simple), 0.4–1 mm, sparsely pubescent or glabrous distally.

Basal leaves

blade broadly oblanceolate to obovate, 7–25(–50) mm wide, margins usually repand to coarsely dentate, rarely entire, not ciliate, surfaces moderately to densely pubescent, trichomes short- to long-stalked, 3–6-rayed, 0.2–0.5 mm.

blade oblanceolate, 3–7(–11) mm wide, margins entire or dentate, ciliate proximally, trichomes to 1 mm, surfaces densely pubescent, trichomes long-stalked, 2–4-rayed, 0.4–0.8 mm.

Cauline leaves

(3–)8–30, not concealing stem;

blade auricles absent, surfaces of distalmost leaves sparsely pubescent.

5–15(–28), often concealing stem proximally;

blade auricles 0.5–3(–5) mm, surfaces of distalmost leaves pubescent.

Racemes

7–25-flowered, sometimes branched.

7–20(–30)-flowered, usually unbranched.

Flowers

erect to ascending at anthesis;

sepals glabrous or sparsely pubescent;

petals white, 3.5–6 × 0.8–1 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals usually purple (rarely lavender), 7–12 × 2–4 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

erect to divaricate-ascending, straight, 3–10 mm, glabrous or pubescent, trichomes spreading, usually 2–5-rayed, rarely simple.

ascending to divaricate-ascending, straight, 3–25 mm, pubescent or glabrous, trichomes spreading, simple and 2-rayed.

Fruits

divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (3.5–)5–13.5 cm × 2.5–4 mm;

valves glabrous or pubescent throughout;

ovules 34–50 per ovary;

style 0.5–1.5 mm.

ascending to divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.5–10 cm × 1.5–2.2 mm;

valves usually glabrous, rarely sparsely pubescent;

ovules 48–96 per ovary;

style 0.05–0.3 mm.

Seeds

uniseriate, 2.5–5(–6) × 2–3(–3.5) mm;

wing continuous, 0.7–1.8 mm wide.

uniseriate, 1.2–1.7 × 1–1.2 mm;

wing continuous, 0.1–0.2 mm wide distally.

2n

= 14.

Boechera repanda

Boechera breweri

Phenology Flowering Jun–Jul.
Habitat Rock outcrops, talus, and gravelly soil in meadows and open pine forests
Elevation 1600-3500 m (5200-11500 ft)
Distribution
from FNA
CA; NV
[WildflowerSearch map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera repanda is a sexual diploid that is distinguished from all other western North American species by its wide (10–50 mm), repand to coarsely dentate basal leaves. The form recognized as var. greenei appears to be a growth response to ecologically marginal habitats. Boechera repanda occurs sporadically in the Sierra Nevada and Transverse Ranges of California, and barely extends into the Carson Range in western Nevada.

Arabis inamoena Greene (1911), not Greene (1908) is an illegitimate name, sometimes found in synonymy with Boechera repanda.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Relatively few other sexual diploid species of Boechera co-occur with B. breweri, and the only currently recognized hybrid involving this taxon is B. rubicundula (= B. arcuata × B. breweri; M. D. Windham and I. A. Al-Shehbaz 2007). Boechera breweri appears to be related to B. sparsiflora, and some specimens of subsp. shastaensis can be difficult to separate from that species. The two subspecies appear to intergrade on the western side of the Sacramento Valley from Tehama to Solano counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems 0.6-2 dm, pubescent distally; fruiting pedicels 3-8(-12) mm, pubescent; fruits 3.5-7.5 cm.
subsp. breweri
1. Stems 1.8-3.5(-4.5) dm, glabrous distally; fruiting pedicels (7-)10-25 mm, usually glabrous; fruits (4.5-)7-10 cm.
subsp. shastaensis
Source FNA vol. 7, p. 402. FNA vol. 7, p. 367.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Subordinate taxa
B. breweri subsp. breweri, B. breweri subsp. shastaensis
Synonyms Arabis repanda, Arabis repanda var. greenei Arabis breweri, Arabis breweri var. figularis, Arabis epilobioides
Name authority (S. Watson) Al-Shehbaz: Novon 13: 388. (2003) (S. Watson) Al-Shehbaz: Novon 13: 384. (2003)
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