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Greene's rockcress, Yosemite rockcress

Habit Biennials or perennials; short-lived; apomictic; caudex present or absent. Perennials; short- to long-lived; sexual; caudex usually not woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4.5 dm, densely pubescent proximally, trichomes sessile, 2–4-rayed, 0.15–0.5 mm, glabrous distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–9 dm, densely pubescent proximally, trichomes short- to long-stalked, 2–6-rayed, 0.2–0.5 mm, rarely mixed with simple ones, to 1.5 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade oblanceolate, 5–15 mm wide, margins denticulate, ciliate proximally, trichomes (simple), to 1 mm, surfaces moderately pubescent, trichomes subsessile, (2- or) 3–7-rayed, 0.1–0.3 mm.

blade broadly oblanceolate to obovate, 7–25(–50) mm wide, margins usually repand to coarsely dentate, rarely entire, not ciliate, surfaces moderately to densely pubescent, trichomes short- to long-stalked, 3–6-rayed, 0.2–0.5 mm.

Cauline leaves

4–15, not concealing stem;

blade auricles 1–3.5 mm, surfaces of distalmost leaves glabrous.

(3–)8–30, not concealing stem;

blade auricles absent, surfaces of distalmost leaves sparsely pubescent.

Racemes

11–41-flowered, usually unbranched.

7–25-flowered, sometimes branched.

Flowers

divaricate at anthesis;

sepals pubescent;

petals white, 6–7 × 1–2 mm, glabrous;

pollen spheroid.

erect to ascending at anthesis;

sepals glabrous or sparsely pubescent;

petals white, 3.5–6 × 0.8–1 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

horizontal to slightly descending, curved to straight, 3–8(–14) mm, glabrous or with some subappressed, branched trichomes.

erect to divaricate-ascending, straight, 3–10 mm, glabrous or pubescent, trichomes spreading, usually 2–5-rayed, rarely simple.

Fruits

horizontal to slightly descending, not appressed to rachis, secund, straight, edges parallel, 3–6 cm × 1.5–2 mm;

valves glabrous;

ovules 56–94 per ovary.

divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (3.5–)5–13.5 cm × 2.5–4 mm;

valves glabrous or pubescent throughout;

ovules 34–50 per ovary;

style 0.5–1.5 mm.

Seeds

uniseriate, 1.2–1.5 × 1–1.3 mm;

wing continuous, 0.1–0.15 mm wide.

uniseriate, 2.5–5(–6) × 2–3(–3.5) mm;

wing continuous, 0.7–1.8 mm wide.

2n

= 14.

Boechera quebecensis

Boechera repanda

Phenology Flowering Jun–Jul. Flowering Jun–Jul.
Habitat Calcareous rock outcrops and talus slopes Rock outcrops, talus, and gravelly soil in meadows and open pine forests
Elevation 0-300 m (0-1000 ft) 1600-3500 m (5200-11500 ft)
Distribution
from FNA
QC
[BONAP county map]
from FNA
CA; NV
[WildflowerSearch map]
Discussion

Morphological evidence suggests that Boechera quebecensis is an apomictic species that arose through hybridization between B. holboellii and B. stricta; it is most likely to be confused with B. grahamii (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera repanda is a sexual diploid that is distinguished from all other western North American species by its wide (10–50 mm), repand to coarsely dentate basal leaves. The form recognized as var. greenei appears to be a growth response to ecologically marginal habitats. Boechera repanda occurs sporadically in the Sierra Nevada and Transverse Ranges of California, and barely extends into the Carson Range in western Nevada.

Arabis inamoena Greene (1911), not Greene (1908) is an illegitimate name, sometimes found in synonymy with Boechera repanda.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 401. FNA vol. 7, p. 402.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis divaricarpa var. dechamplainii Arabis repanda, Arabis repanda var. greenei
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 246. (2007) (S. Watson) Al-Shehbaz: Novon 13: 388. (2003)
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