Boechera quebecensis |
Boechera formosa |
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beautiful rockcress |
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Habit | Biennials or perennials; short-lived; apomictic; caudex present or absent. | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4.5 dm, densely pubescent proximally, trichomes sessile, 2–4-rayed, 0.15–0.5 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of leaf tuft, above ground surface on woody base, 2–5.5 dm, densely pubescent proximally, trichomes short-stalked, 4–7-rayed, 0.1–0.4 mm, similarly pubescent distally. |
Basal leaves | blade oblanceolate, 5–15 mm wide, margins denticulate, ciliate proximally, trichomes (simple), to 1 mm, surfaces moderately pubescent, trichomes subsessile, (2- or) 3–7-rayed, 0.1–0.3 mm. |
blade linear to linear-oblanceolate, 2–3(–4) mm wide, margins entire, sometimes ciliate near petiole base, trichomes (simple or 2-rayed), to 0.8 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
Cauline leaves | 4–15, not concealing stem; blade auricles 1–3.5 mm, surfaces of distalmost leaves glabrous. |
7–18, concealing stem proximally; blade auricles absent, surfaces of distalmost leaves densely pubescent. |
Racemes | 11–41-flowered, usually unbranched. |
6–26-flowered, usually unbranched. |
Flowers | divaricate at anthesis; sepals pubescent; petals white, 6–7 × 1–2 mm, glabrous; pollen spheroid. |
divaricate at anthesis; sepals pubescent; petals white to pale lavender, 8–18 × 2.5–4.4(–5.5) mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | horizontal to slightly descending, curved to straight, 3–8(–14) mm, glabrous or with some subappressed, branched trichomes. |
horizontal to descending, usually recurved, 5–10(–20) mm, pubescent, trichomes appressed, branched. |
Fruits | horizontal to slightly descending, not appressed to rachis, secund, straight, edges parallel, 3–6 cm × 1.5–2 mm; valves glabrous; ovules 56–94 per ovary. |
divaricate-descending to reflexed, usually not appressed to rachis, rarely somewhat secund, straight, edges parallel, 4.3–7 cm × 1.6–3 mm; valves pubescent throughout; ovules 72–98 per ovary; style ca. 0.1 mm. |
Seeds | uniseriate, 1.2–1.5 × 1–1.3 mm; wing continuous, 0.1–0.15 mm wide. |
biseriate, 1.2–1.6 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 14. |
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Boechera quebecensis |
Boechera formosa |
|
Phenology | Flowering Jun–Jul. | Flowering Apr–Jun. |
Habitat | Calcareous rock outcrops and talus slopes | Rocky slopes and sandy soil in blackbrush, sagebrush, and pinyon-juniper communities |
Elevation | 0-300 m (0-1000 ft) | 1300-1900 m (4300-6200 ft) |
Distribution |
QC |
AZ; CO; NM; UT; WY
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Discussion | Morphological evidence suggests that Boechera quebecensis is an apomictic species that arose through hybridization between B. holboellii and B. stricta; it is most likely to be confused with B. grahamii (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although Boechera formosa is usually subsumed within Arabis (Boechera) pulchra, the two species are easily separated. Boechera formosa has white to lavender (versus mostly purple) petals, fruiting pedicels that are not abruptly recurved at the base, narrower (1.6–3 versus 2.5–4 mm wide), mostly non-appressed fruits, and narrower (1–1.2 versus 1.5–2.2 mm wide) seeds. The two taxa are separated by more than 500 km, with B. formosa found only on the Colorado Plateau and B. pulchra, in the strict sense, restricted to southern California, western Nevada, and northwestern Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 401. | FNA vol. 7, p. 379. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis divaricarpa var. dechamplainii | Arabis formosa, Arabis pulchra var. pallens, B. pulchra subsp. pallens, B. pulchra var. pallens |
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 246. (2007) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 68. (2006) |
Web links |