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Constance's Rock cress

Habit Biennials or perennials; short-lived; apomictic; caudex present or absent. Perennials; long-lived; sexual; caudex woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4.5 dm, densely pubescent proximally, trichomes sessile, 2–4-rayed, 0.15–0.5 mm, glabrous distally.

usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 1.2–3 dm, glabrous throughout.

Basal leaves

blade oblanceolate, 5–15 mm wide, margins denticulate, ciliate proximally, trichomes (simple), to 1 mm, surfaces moderately pubescent, trichomes subsessile, (2- or) 3–7-rayed, 0.1–0.3 mm.

blade narrowly oblanceolate, 1.5–4 mm wide, margins entire, ciliate, trichomes (simple, mixed with fewer stalked, 2-rayed), 0.3–0.8 mm, surfaces glabrous.

Cauline leaves

4–15, not concealing stem;

blade auricles 1–3.5 mm, surfaces of distalmost leaves glabrous.

6–12, not concealing stem;

blade auricles absent, surfaces of distalmost leaves glabrous.

Racemes

11–41-flowered, usually unbranched.

5–15-flowered, unbranched.

Flowers

divaricate at anthesis;

sepals pubescent;

petals white, 6–7 × 1–2 mm, glabrous;

pollen spheroid.

divaricate-ascending at anthesis;

sepals glabrous;

petals creamy white, 6–8 × 1.5–2 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

horizontal to slightly descending, curved to straight, 3–8(–14) mm, glabrous or with some subappressed, branched trichomes.

arched, strongly recurved proximally, 4–12 mm, glabrous.

Fruits

horizontal to slightly descending, not appressed to rachis, secund, straight, edges parallel, 3–6 cm × 1.5–2 mm;

valves glabrous;

ovules 56–94 per ovary.

pendent, not appressed to rachis, usually secund, straight or slightly curved, edges often somewhat undulate (not parallel), 4–7.5 cm × 3–3.5 mm;

valves glabrous;

ovules 18–28 per ovary;

style 2.5–5 mm.

Seeds

uniseriate, 1.2–1.5 × 1–1.3 mm;

wing continuous, 0.1–0.15 mm wide.

uniseriate, 3–4 × 2.5–3 mm;

wing continuous, 0.5–1 mm wide.

2n

= 14.

Boechera quebecensis

Boechera constancei

Phenology Flowering Jun–Jul. Flowering May.
Habitat Calcareous rock outcrops and talus slopes Steep ridges, serpentine slopes
Elevation 0-300 m (0-1000 ft) 1200-1900 m (3900-6200 ft)
Distribution
from FNA
QC
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Morphological evidence suggests that Boechera quebecensis is an apomictic species that arose through hybridization between B. holboellii and B. stricta; it is most likely to be confused with B. grahamii (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera constancei was originally treated as a variety of Arabis (Boechera) suffrutescens; it is easily separated from that species by its non-auriculate cauline leaf blades, longer (6–8 versus 4.5–6 mm) petals that are creamy white (versus purple or rose-tipped), much longer style (2.5–5 versus 0.4–1.2 mm), and exerted anthers. This distinctive sexual diploid is thus far known only from the west slope of the Sierra Nevada in Plumas County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 401. FNA vol. 7, p. 371.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis divaricarpa var. dechamplainii Arabis constancei, Arabis suffrutescens var. perstylosa
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 246. (2007) (Rollins) Al-Shehbaz: Novon 13: 384. (2003)
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