Boechera pusilla |
Boechera inyoensis |
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Inyo rockcress |
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Habit | Perennials; long-lived; (cespitose); apomictic; caudex often woody. | Perennials; short-lived; apomictic; caudex usually not woody. |
Stems | usually 2–6 per caudex branch, arising from margin of rosette near ground surface, 0.5–2 dm, glabrous or sparsely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 0.2 mm, glabrous distally. |
usually 1 per caudex, arising from center of rosette near ground surface, (1–)2.5–6.5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
Basal leaves | blade linear-oblanceolate, 1–2.5 mm wide, margins entire, ciliate along petiole, trichomes (simple), 0.4–0.7 mm, surfaces usually sparsely pubescent, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.1–0.4 mm. |
blade oblanceolate, 1–4(–8) mm wide, margins entire, ciliate along petiole and proximally, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 3–10-rayed, 0.2–0.7 mm. |
Cauline leaves | 3–5, not concealing stem; blade auricles 0–0.2 mm, surfaces of distalmost leaves usually glabrous or, rarely, margins sparsely ciliate. |
(7–)12–35, concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves densely pubescent. |
Racemes | 6–13-flowered, unbranched. |
10–65-flowered, usually unbranched. |
Flowers | divaricate-ascending at anthesis; sepals glabrous or sparsely pubescent, trichomes spreading, 2-rayed; petals white to lavender, 4–5 × 1.5–1.8 mm, glabrous; pollen spheroid. |
ascending at anthesis; sepals pubescent; petals lavender to purplish, 5–8 × 1.2–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | horizontal to divaricate-descending, straight or slightly curved downward, 2–5 mm, glabrous. |
divaricate-ascending to horizontal, straight or slightly curved downward, 5–15 mm, sparsely pubescent, trichomes appressed, branched. |
Fruits | horizontal or divaricate-descending, not appressed to rachis, secund, straight, edges parallel, 1.6–3.2 cm × 1.5–2 mm; valves glabrous; ovules 20–32 per ovary; style 0.1–0.4 mm. |
divaricate-ascending to widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.7–6.5 cm × 1.5–2.2 mm; valves glabrous; ovules 74–134 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 1.2–1.5 × 0.8–0.9 mm; not winged or with distal wing 0.05–0.1 mm wide. |
usually sub-biseriate, rarely uniseriate, 1.7–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 21. |
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Boechera pusilla |
Boechera inyoensis |
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Phenology | Flowering May–Jun. | Flowering Apr–Jun. |
Habitat | Cracks and crevices of granitic rock outcrops | Limestone and volcanic rock outcrops and clay soils in desert scrub and pinyon-juniper woodlands |
Elevation | 2400-2500 m (7900-8200 ft) | 1400-2400 m (4600-7900 ft) |
Distribution |
WY |
CA; NV; UT
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Discussion | Of conservation concern. Morphological evidence suggests that Boechera pusilla is an apomictic species that arose through hybridization between B. lemmonii and B. pendulina; it is known only from the type locality in southwestern Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Some plants here assigned to Boechera inyoensis were treated by R. C. Rollins (1993) and N. H. Holmgren (2005b) as Arabis (or B.) beckwithii. Closer examination of the type specimens of A. beckwithii reveals that they are conspecific with B. puberula, a sexual diploid widespread in the Great Basin of western North America. Boechera inyoensis is an apomictic triploid that clearly contains at least one genome derived from B. shockleyi (M. D. Windham and I. A. Al-Shehbaz 2007b); the other parent has not been determined. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 400. | FNA vol. 7, p. 385. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis pusilla | Arabis inyoensis, Arabis holboellii var. derensis, B. selbyi var. inyoensis |
Name authority | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) | (Rollins) Al-Shehbaz: Novon 13: 386. (2003) |
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