Boechera pulchra |
Boechera pendulocarpa |
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beautiful rockcress |
danglepod rockcress, drop-seed rock-cress, earless suncress, Holboell's rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody. | Perennials; short- to long-lived; sexual; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of leaf tufts, elevated above ground surface on woody base, (1.5–)3–7.5 dm, densely pubescent proximally, trichomes short-stalked, usually 4–7-rayed, 0.1–0.3 mm, similarly pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–3 dm, densely pubescent proximally, trichomes simple and stalked, 2–4-rayed, 0.1–0.4 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade linear to linear-oblanceolate, 1–3 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 4–9-rayed, 0.1–0.3 mm. |
blade narrowly oblanceolate, 1.5–4(–5) mm wide, margins entire, often ciliate along petiole, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.08–0.2 mm. |
Cauline leaves | 10–30, often concealing stem proximally; blade auricles absent or, rarely, to 0.5 mm, surfaces of distalmost leaves pubescent. |
6–17, sometimes concealing stem proximally; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 8–25-flowered, usually unbranched. |
4–11-flowered, unbranched. |
Flowers | descending at anthesis; sepals pubescent; petals usually purple, rarely white, 9–16 × 2–4(–5) mm, sparsely pubescent or glabrous (trichomes scattered abaxially); pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white or lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 8–16 mm, pubescent, trichomes appressed, branched. |
arched, gently recurved, 3–8(–12) mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | strongly reflexed, often appressed to rachis, sometimes somewhat secund, straight, edges parallel, 3.3–8 cm × 2.5–4 mm; valves pubescent throughout; ovules 68–106 per ovary; style 0.1–0.3 mm. |
pendent, not appressed to rachis, not secund, usually straight, edges parallel, (2–)2.5–3.8 cm × 1.5–2.2 mm; valves glabrous; ovules 66–92 per ovary; style 0.3–0.5 mm. |
Seeds | biseriate, 1.7–2.8 × 1.5–2.2 mm; wing continuous, 0.25–0.65 mm wide. |
sub-biseriate, 1–1.2 × 0.7–0.9 mm; wing distal, 0.05–0.1 mm wide. |
2n | = 14. |
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Boechera pulchra |
Boechera pendulocarpa |
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Phenology | Flowering Mar–Jun. | Flowering Apr–Jul. |
Habitat | Rocky, gravelly or sandy slopes in chaparral, sagebrush and desert scrub communities | Rock outcrops and gravelly slopes in sagebrush, open conifer forests, subalpine meadows |
Elevation | 800-2800 m (2600-9200 ft) | 1000-3300 m (3300-10800 ft) |
Distribution |
CA; NV; Mexico (Baja California)
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CA; ID; MT; NV; OR; UT; WY; BC; YT
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Discussion | Boechera pulchra is a distinctive diploid separated from related sexual species (B. formosa and B. lincolnensis) by abruptly recurved fruiting pedicels and purple flowers, and from apomictic hybrids (e.g., B. xylopoda) by having fruits densely pubescent throughout. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993), B. pendulocarpa is easily distinguished from that species by having simple and 2–4-rayed (versus 4–8-rayed) trichomes proximally on stems, cauline leaves without auricles, fruiting pedicels gently (versus sharply) recurved, and shorter (2–)2.5–3.8 (versus 3.5–6.5) cm, non-secund fruits. The two taxa have allopatric distributions, with B. pendulocarpa found in the mountains of western North America and B. holboellii apparently confined to Greenland. Recent use of the name A. (Boechera) exilis for this taxon (e.g., G. A. Mulligan 1996; R. D. Dorn 2001; N. H. Holmgren 2005b) is based on misinterpretation of the type (M. D. Windham and I. A. Al-Shehbaz 2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 400. | FNA vol. 7, p. 395. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis pulchra | Arabis pendulocarpa, Arabis holboellii var. pendulocarpa, B. holboellii var. pendulocarpa |
Name authority | (M. E. Jones ex S. Watson) W. A. Weber: Phytologia 51: 370. (1982) | (A. Nelson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 77. (2006) |
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