Boechera pulchra |
Boechera holboellii |
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beautiful rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody. | Biennials or perennials; short-lived; apomictic or sexual; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of leaf tufts, elevated above ground surface on woody base, (1.5–)3–7.5 dm, densely pubescent proximally, trichomes short-stalked, usually 4–7-rayed, 0.1–0.3 mm, similarly pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6.7 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade linear to linear-oblanceolate, 1–3 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 4–9-rayed, 0.1–0.3 mm. |
blade oblanceolate, 2–5 mm wide, margins entire, sometimes ciliate near petiole base, trichomes (usually simple), to 0.7 mm, surfaces sparsely to moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
Cauline leaves | 10–30, often concealing stem proximally; blade auricles absent or, rarely, to 0.5 mm, surfaces of distalmost leaves pubescent. |
7–40, often concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 8–25-flowered, usually unbranched. |
10–60-flowered, usually unbranched. |
Flowers | descending at anthesis; sepals pubescent; petals usually purple, rarely white, 9–16 × 2–4(–5) mm, sparsely pubescent or glabrous (trichomes scattered abaxially); pollen ellipsoid. |
divaricate to pendent at anthesis; sepals pubescent; petals white, 4–8 × 1–2 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 8–16 mm, pubescent, trichomes appressed, branched. |
descending to reflexed, most abruptly recurved near base, 5–10 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | strongly reflexed, often appressed to rachis, sometimes somewhat secund, straight, edges parallel, 3.3–8 cm × 2.5–4 mm; valves pubescent throughout; ovules 68–106 per ovary; style 0.1–0.3 mm. |
descending to reflexed, not appressed to rachis, strongly secund, often curved, edges parallel, 0.35–0.65 cm × (2–)2.2–2.5 mm; valves glabrous; ovules 80–102 per ovary. |
Seeds | biseriate, 1.7–2.8 × 1.5–2.2 mm; wing continuous, 0.25–0.65 mm wide. |
uniseriate to sub-biseriate, 1.4–1.7 × 1–1.2 mm; wing continuous, to 0.2 mm wide. |
2n | = 14, 21. |
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Boechera pulchra |
Boechera holboellii |
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Phenology | Flowering Mar–Jun. | Flowering Jul–Aug. |
Habitat | Rocky, gravelly or sandy slopes in chaparral, sagebrush and desert scrub communities | Rocky slopes and gravelly soil near coast |
Elevation | 800-2800 m (2600-9200 ft) | 100-300 m (300-1000 ft) |
Distribution |
CA; NV; Mexico (Baja California)
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Greenland |
Discussion | Boechera pulchra is a distinctive diploid separated from related sexual species (B. formosa and B. lincolnensis) by abruptly recurved fruiting pedicels and purple flowers, and from apomictic hybrids (e.g., B. xylopoda) by having fruits densely pubescent throughout. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Both sexual diploid and apomictic triploid collections of Boechera holboellii are known; available data suggest that the triploid is an autopolyploid derivative of the sexual diploid but further study is needed. In the narrow circumscription adopted here, it appears to be restricted to Greenland (see discussion of B. quebecensis in M. D. Windham and I. A. Al-Shehbaz 2007b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 400. | FNA vol. 7, p. 384. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis pulchra | Arabis holboellii, Arabis holboellii var. tenuis, B. tenuis |
Name authority | (M. E. Jones ex S. Watson) W. A. Weber: Phytologia 51: 370. (1982) | (Hornemann) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) |
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