Boechera pulchra |
Boechera covillei |
|
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beautiful rockcress |
Coville's rock-cress |
|
Habit | Perennials; long-lived; sexual; caudex woody. | Perennials; long-lived; apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of leaf tufts, elevated above ground surface on woody base, (1.5–)3–7.5 dm, densely pubescent proximally, trichomes short-stalked, usually 4–7-rayed, 0.1–0.3 mm, similarly pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.5–2.5 dm, glabrous throughout. |
Basal leaves | blade linear to linear-oblanceolate, 1–3 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 4–9-rayed, 0.1–0.3 mm. |
blade oblanceolate, 1.5–5 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.4 mm, and distally, trichomes short-stalked, 2 or 3 (or 4)-rayed, 0.15–0.4 mm, surfaces glabrous. |
Cauline leaves | 10–30, often concealing stem proximally; blade auricles absent or, rarely, to 0.5 mm, surfaces of distalmost leaves pubescent. |
2–7, not concealing stem; blade with auricles 0.2–1 mm, auricles rarely absent, surfaces of distalmost leaves glabrous. |
Racemes | 8–25-flowered, usually unbranched. |
2–9-flowered, unbranched. |
Flowers | descending at anthesis; sepals pubescent; petals usually purple, rarely white, 9–16 × 2–4(–5) mm, sparsely pubescent or glabrous (trichomes scattered abaxially); pollen ellipsoid. |
ascending at anthesis; sepals glabrous; petals lavender to purplish, 5–6 × 1–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 8–16 mm, pubescent, trichomes appressed, branched. |
ascending to suberect, straight, 6–20 mm, glabrous. |
Fruits | strongly reflexed, often appressed to rachis, sometimes somewhat secund, straight, edges parallel, 3.3–8 cm × 2.5–4 mm; valves pubescent throughout; ovules 68–106 per ovary; style 0.1–0.3 mm. |
ascending to suberect, rarely appressed to rachis, not secund, straight, edges slightly undulate (rarely parallel), 3.5–5 cm × 3–5 mm; valves glabrous; ovules 18–30 per ovary; style 0.2–1 mm. |
Seeds | biseriate, 1.7–2.8 × 1.5–2.2 mm; wing continuous, 0.25–0.65 mm wide. |
uniseriate, 4–5 × 3–4 mm; wing continuous, 1–1.5 mm wide. |
Boechera pulchra |
Boechera covillei |
|
Phenology | Flowering Mar–Jun. | Flowering Jul–Aug. |
Habitat | Rocky, gravelly or sandy slopes in chaparral, sagebrush and desert scrub communities | Rocky slopes in alpine meadows and open coniferous forests |
Elevation | 800-2800 m (2600-9200 ft) | 2200-3500 m (7200-11500 ft) |
Distribution |
CA; NV; Mexico (Baja California)
|
CA; NV; OR |
Discussion | Boechera pulchra is a distinctive diploid separated from related sexual species (B. formosa and B. lincolnensis) by abruptly recurved fruiting pedicels and purple flowers, and from apomictic hybrids (e.g., B. xylopoda) by having fruits densely pubescent throughout. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera covillei is an apomictic species that arose through hybridization between B. howellii and B. lyallii (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 400. | FNA vol. 7, p. 371. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis pulchra | Arabis covillei, Arabis leibergii |
Name authority | (M. E. Jones ex S. Watson) W. A. Weber: Phytologia 51: 370. (1982) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 237. (2007) |
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