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Blue Mountain rockcress, hoary rock-cress, silver rockcress

dangling suncress, reflexed rock-cress, secund rock-cress

Habit Perennials; short-lived; sexual; caudex not woody. Biennials or perennials; short-lived; sexual; caudex present or absent.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3(–0.5) mm, similarly pubescent distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–7(–10.5) dm, densely pubescent proximally, trichomes short-stalked, 2–8-rayed, 0.1–0.2 mm, sparsely pubescent distally.

Basal leaves

blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm.

blade oblanceolate, 2–7 mm wide, margins entire or shallowly dentate, sometimes ciliate at petiole base, surfaces densely pubescent, trichomes short-stalked, 5–10-rayed, 0.05–0.2 mm.

Cauline leaves

7–45(–65), concealing stem proximally;

blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent.

15–40, concealing stem proximally;

blade auricles 0.5–2.5 mm or, rarely, absent, surfaces of distalmost leaves pubescent.

Racemes

10–40(–64)-flowered, usually unbranched.

15–80(–140)-flowered, usually unbranched.

Flowers

ascending to descending at anthesis;

sepals pubescent;

petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous;

pollen ellipsoid.

pendent at anthesis;

sepals pubescent;

petals white to lavender, 4–8 × 0.8–2.2 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched.

reflexed, abruptly recurved at base, otherwise usually straight, 7–12(–18) mm, pubescent, trichomes appressed, branched.

Fruits

closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm;

valves pubescent throughout;

ovules 38–64 per ovary;

style 0.05–0.1 mm.

strongly reflexed, usually appressed to rachis, sometimes somewhat secund, straight, edges parallel, 3.5–9 cm × 0.9–1.8 mm;

valves glabrous or sparsely pubescent throughout;

ovules 60–116 per ovary;

style 0.5–2 mm.

Seeds

uniseriate, 1.4–1.8 × 1–1.4 mm;

wing continuous, 0.1–0.3 mm wide.

uniseriate, 1–1.4 × 0.9–1.2 mm;

wing continuous, to 0.1 mm wide.

2n

= 14.

= 14.

Boechera puberula

Boechera retrofracta

Phenology Flowering Apr–Jul. Flowering Apr–Aug.
Habitat Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities Rock outcrops, open hillsides, gravel bars and sandy banks in grassland, sagebrush, oak woodlands, and open conifer forests
Elevation 1300-2900 m (4300-9500 ft) 300-3300 m (1000-10800 ft)
Distribution
from FNA
CA; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; ID; MI; MT; NV; OR; UT; WA; WY; AB; BC; MB; NT; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996; N. H. Holmgren 2005b), B. retrofracta is easily distinguished from that species by having narrower (0.9–1.8 versus 2–2.5 mm), mostly non-secund fruits that are almost always appressed to rachises. The two taxa have allopatric distributions, with B. retrofracta found on the North American continent (mostly west of the Great Plains) and B. holboellii apparently confined to Greenland. Boechera retrofracta has formed hybrids with at least 12 other species. Besides differing in macromorphological characters, all those hybrids are distinct from B. retrofracta in the strict sense in having wider (20–30 versus 13–16 µm), spheroid pollen grains with asymmetric colpi.

Arabis kochii Blankinship is an illegitimate name, sometimes found in synonymy with Boechera retrofracta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 399. FNA vol. 7, p. 402.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis puberula, Arabis arida, Arabis beckwithii, Arabis lignipes var. impar, Arabis sabulosa, Arabis subpinnatifida var. beckwithii, Arabis subpinnatifida var. impar, B. beckwithii Arabis retrofracta, Arabis arcuata var. secunda, Arabis exilis, Arabis holboellii var. retrofracta, Arabis holboellii var. secunda, Arabis lignipes, Arabis retrofracta var. multicaulis, Arabis secunda, Arabis sparsiflora subsp. secunda, Arabis tenuis, B. exilis, B. holboellii var. secunda
Name authority (Nuttall) Dorn: Brittonia, 55: 3. (2003) (Graham) Á. Löve & D. Löve: Taxon 31: 125. (1982)
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