Boechera puberula |
Boechera gracilipes |
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Blue Mountain rockcress, hoary rock-cress, silver rockcress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Biennials or perennials; short-lived; sexual; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3(–0.5) mm, similarly pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, rarely arising laterally proximal to sterile shoots, 2.5–8.5 dm, densely pubescent proximally, trichomes simple or spurred, 0.9–1.5 mm, glabrous distally. |
Basal leaves | blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm. |
blade oblanceolate, 5–12 mm wide, margins shallowly dentate, sometimes ciliate near petiole base, surfaces sparsely to densely pubescent, trichomes short- to long-stalked, 2- or 3-rayed, usually 0.3–0.6 mm. |
Cauline leaves | 7–45(–65), concealing stem proximally; blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent. |
30–65, often concealing stem proximally; blade auricles 0.8–3 mm, surfaces of distalmost leaves usually glabrous. |
Racemes | 10–40(–64)-flowered, usually unbranched. |
(12–)15–50-flowered, usually unbranched. |
Flowers | ascending to descending at anthesis; sepals pubescent; petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white to pale lavender, 6–9 × 0.6–1 mm; valves glabrous; pollen ellipsoid. |
Fruiting pedicels | pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending or horizontal, recurved, (15–)20–47 mm, glabrous. |
Fruits | closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm; valves pubescent throughout; ovules 38–64 per ovary; style 0.05–0.1 mm. |
widely pendent, not appressed to rachis, not secund, curved, edges parallel, (2.5–)3–7.5 cm × 1.5–2.8 mm, glabrous; ovules 130–210 per ovary; style 0.2–0.5 mm. |
Seeds | uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.3 mm wide. |
biseriate, 1.2–1.4 × 0.7–0.9 mm; wing continuous or distal, to 0.2 mm wide. |
2n | = 14. |
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Boechera puberula |
Boechera gracilipes |
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Phenology | Flowering Apr–Jul. | Flowering Apr–Jun. |
Habitat | Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities | Basalt, limestone, and sandy soils in ponderosa pine forests and pinyon-juniper woodlands |
Elevation | 1300-2900 m (4300-9500 ft) | 1700-2300 m (5600-7500 ft) |
Distribution |
CA; NV; OR; UT
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AZ; NV; UT |
Discussion | Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fruiting pedicel length, considered diagnostic by R. C. Rollins (1993) and N. H. Holmgren (2005b), occasionally fails to separate Boechera gracilipes from the closely-related B. fendleri. The two are consistently distinguished by trichome characters. In B. gracilipes, basal leaves usually lack prominent cilia and surfaces are persistently pubescent with at least some 3-rayed trichomes. Also, stems are rather densely pilose proximally, with the largest trichomes more than 0.9 mm. By contrast, basal leaves of B. fendleri always have prominent cilia, surfaces are often glabrescent, and 3-rayed trichomes usually are rare or absent. Stems of the latter are proximally hirsute to hispid, with the largest trichomes less than 0.9 mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 399. | FNA vol. 7, p. 381. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis puberula, Arabis arida, Arabis beckwithii, Arabis lignipes var. impar, Arabis sabulosa, Arabis subpinnatifida var. beckwithii, Arabis subpinnatifida var. impar, B. beckwithii | Arabis gracilipes, Arabis arcuata var. longipes, Arabis perennans var. longipes |
Name authority | (Nuttall) Dorn: Brittonia, 55: 3. (2003) | (Greene) Dorn: Brittonia 55: 3. (2003) |
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