Boechera puberula |
Boechera fernaldiana |
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Blue Mountain rockcress, hoary rock-cress, silver rockcress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; long-lived; (cespitose); sexual; caudex somewhat woody (sometimes with persistent, crowded leaf bases). | ||||
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3(–0.5) mm, similarly pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, near ground surface or slightly elevated on woody base, 1–3.8 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.04–0.1 mm, sometimes mixed with stalked, 2-rayed ones, to 0.5 mm, usually sparsely pubescent distally. |
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Basal leaves | blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm. |
blade narrowly oblanceolate, 1–4 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.04–0.1 mm. |
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Cauline leaves | 7–45(–65), concealing stem proximally; blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent. |
5–10, not concealing stem; blade auricles 0.3–2 mm, surfaces of distalmost leaves usually sparsely pubescent. |
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Racemes | 10–40(–64)-flowered, usually unbranched. |
10–20-flowered, usually unbranched. |
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Flowers | ascending to descending at anthesis; sepals pubescent; petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals (purplish or greenish) glabrous or sparsely or moderately pubescent; petals white or purple to lavender, 7–12 × 1.5–4 mm, glabrous; pollen ellipsoid. |
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Fruiting pedicels | pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending, straight, 5–20 mm, sparsely pubescent, trichomes appressed, branched. |
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Fruits | closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm; valves pubescent throughout; ovules 38–64 per ovary; style 0.05–0.1 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight or curved, edges parallel, 3.5–6.5(–7.5) cm × 1–1.6 mm; valves glabrous; ovules 30–72 per ovary; style 0.1–1 mm. |
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Seeds | uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.3 mm wide. |
uniseriate, 1–1.2 × 0.8–1 mm; wing continuous, ca. 0.1 mm wide. |
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2n | = 14. |
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Boechera puberula |
Boechera fernaldiana |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities | |||||
Elevation | 1300-2900 m (4300-9500 ft) | |||||
Distribution |
CA; NV; OR; UT
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CO; NV; UT |
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Discussion | Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Taxa treated here as subspecies of Boechera fernaldiana share many morphological traits, and most recent authors (R. C. Rollins 1993; N. H. Holmgren 2005b) did not recognize them as distinct. They are consistently separated by the characters listed below, show some degree of molecular divergence (C. D. Bailey et al., unpubl.), and their ranges are separated by ca. 500 km. They clearly represent genetically isolated population systems that warrant taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 399. | FNA vol. 7, p. 378. | ||||
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Arabis puberula, Arabis arida, Arabis beckwithii, Arabis lignipes var. impar, Arabis sabulosa, Arabis subpinnatifida var. beckwithii, Arabis subpinnatifida var. impar, B. beckwithii | Arabis fernaldiana, Arabis canescens var. stylosa, Arabis fernaldiana var. stylosa | ||||
Name authority | (Nuttall) Dorn: Brittonia, 55: 3. (2003) | (Rollins) W. A. Weber: Phytologia 51: 370. (1982) | ||||
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