Boechera puberula |
Boechera divaricarpa |
|
---|---|---|
Blue Mountain rockcress, hoary rock-cress, silver rockcress |
spreading-pod rock-cress, uplifting suncress |
|
Habit | Perennials; short-lived; sexual; caudex not woody. | Biennials or perennials; short-lived; apomictic; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3(–0.5) mm, similarly pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)3–9 dm, glabrous or pubescent proximally, trichomes sessile, 2–4-rayed (some simple), to 0.7 mm, glabrous distally. |
Basal leaves | blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm. |
blade oblanceolate, 2–10 mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes sessile, 2–6-rayed, 0.1–0.4 mm. |
Cauline leaves | 7–45(–65), concealing stem proximally; blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent. |
(10–)15–56, concealing stem proximally; blade auricles 1–5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 10–40(–64)-flowered, usually unbranched. |
12–40(–65)-flowered, usually unbranched. |
Flowers | ascending to descending at anthesis; sepals pubescent; petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals glabrous or with scattered trichomes; petals usually purple, rarely lavender, 6–9 × 1.5–3 mm, glabrous; pollen spheroid. |
Fruiting pedicels | pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending to horizontal, straight, 5–12 mm, glabrous. |
Fruits | closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm; valves pubescent throughout; ovules 38–64 per ovary; style 0.05–0.1 mm. |
divaricate-ascending to horizontal, not appressed to rachis, not secund, straight, edges parallel, (4.5–)5.5–11 cm × 1.7–2.5 mm; valves glabrous; ovules 114–142 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.3 mm wide. |
uniseriate to sub-biseriate, 1.4–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 14. |
= 21. |
Boechera puberula |
Boechera divaricarpa |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jul. |
Habitat | Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities | Rock outcrops, talus slopes and gravelly hillsides in sagebrush, mountain shrub, and open conifer forests |
Elevation | 1300-2900 m (4300-9500 ft) | 900-2500 m (3000-8200 ft) |
Distribution |
CA; NV; OR; UT
|
CA; ID; MT; NV; UT; WA; WY
|
Discussion | Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The name Arabis (Boechera) divaricarpa has been applied to nearly every hybrid containing a genome derived from B. stricta. This presents a serious barrier to understanding the evolution of Boechera and also is contrary to the International Code of Botanical Nomenclature, because some names usually placed in synonymy (i.e., B. grahamii and B. brachycarpa) have priority at species level (M. D. Windham and I. A. Al-Shehbaz 2007b). To address this problem, we treat the following as distinct species: B. acutina, B. grahamii (= B. brachycarpa of R. D. Dorn 2001), and B. pratincola (all considered synonyms of A. divaricarpa by R. C. Rollins 1993), and B. calderi, B. elkoensis, and B. quebecensis (taxa described after 1993). Detailed comparison among these taxa are provided by Windham and Al-Shehbaz (2007, 2007b). The narrow concept of B. divaricarpa advocated here encompasses apomictic triploid populations containing three distinct genomes, one each derived from B. retrofracta, B. sparsiflora, and B. stricta. If the species is defined more broadly, the name B. grahamii has priority. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 399. | FNA vol. 7, p. 374. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis puberula, Arabis arida, Arabis beckwithii, Arabis lignipes var. impar, Arabis sabulosa, Arabis subpinnatifida var. beckwithii, Arabis subpinnatifida var. impar, B. beckwithii | Arabis divaricarpa, Arabis stokesiae |
Name authority | (Nuttall) Dorn: Brittonia, 55: 3. (2003) | (A. Nelson) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) |
Web links |
|
|