Boechera puberula |
Boechera covillei |
|
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Blue Mountain rockcress, hoary rock-cress, silver rockcress |
Coville's rock-cress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; long-lived; apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3(–0.5) mm, similarly pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.5–2.5 dm, glabrous throughout. |
Basal leaves | blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm. |
blade oblanceolate, 1.5–5 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.4 mm, and distally, trichomes short-stalked, 2 or 3 (or 4)-rayed, 0.15–0.4 mm, surfaces glabrous. |
Cauline leaves | 7–45(–65), concealing stem proximally; blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent. |
2–7, not concealing stem; blade with auricles 0.2–1 mm, auricles rarely absent, surfaces of distalmost leaves glabrous. |
Racemes | 10–40(–64)-flowered, usually unbranched. |
2–9-flowered, unbranched. |
Flowers | ascending to descending at anthesis; sepals pubescent; petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals glabrous; petals lavender to purplish, 5–6 × 1–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched. |
ascending to suberect, straight, 6–20 mm, glabrous. |
Fruits | closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm; valves pubescent throughout; ovules 38–64 per ovary; style 0.05–0.1 mm. |
ascending to suberect, rarely appressed to rachis, not secund, straight, edges slightly undulate (rarely parallel), 3.5–5 cm × 3–5 mm; valves glabrous; ovules 18–30 per ovary; style 0.2–1 mm. |
Seeds | uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.3 mm wide. |
uniseriate, 4–5 × 3–4 mm; wing continuous, 1–1.5 mm wide. |
2n | = 14. |
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Boechera puberula |
Boechera covillei |
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Phenology | Flowering Apr–Jul. | Flowering Jul–Aug. |
Habitat | Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities | Rocky slopes in alpine meadows and open coniferous forests |
Elevation | 1300-2900 m (4300-9500 ft) | 2200-3500 m (7200-11500 ft) |
Distribution |
CA; NV; OR; UT
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CA; NV; OR |
Discussion | Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera covillei is an apomictic species that arose through hybridization between B. howellii and B. lyallii (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 399. | FNA vol. 7, p. 371. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis puberula, Arabis arida, Arabis beckwithii, Arabis lignipes var. impar, Arabis sabulosa, Arabis subpinnatifida var. beckwithii, Arabis subpinnatifida var. impar, B. beckwithii | Arabis covillei, Arabis leibergii |
Name authority | (Nuttall) Dorn: Brittonia, 55: 3. (2003) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 237. (2007) |
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