Boechera puberula |
Boechera californica |
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Blue Mountain rockcress, hoary rock-cress, silver rockcress |
California rockcress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; usually long-lived; apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3(–0.5) mm, similarly pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, (2–)3.5–13 dm, densely pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.3–0.9 mm, moderately to sparsely pubescent distally. |
Basal leaves | blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm. |
blade linear to oblanceolate, 3–10(–13) mm wide, margins entire or denticulate, ciliate along petiole, trichomes to 1.3 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.2–0.5 mm. |
Cauline leaves | 7–45(–65), concealing stem proximally; blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent. |
12–35(–55), often concealing stem proximally; blade auricles 1–5(–6) mm, surfaces of distalmost leaves pubescent. |
Racemes | 10–40(–64)-flowered, usually unbranched. |
30–120-flowered, usually unbranched. |
Flowers | ascending to descending at anthesis; sepals pubescent; petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals usually purple, rarely pinkish, 9–14 × 1.5–3 mm, glabrous or sparsely pubescent (trichomes scattered abaxially); pollen spheroid. |
Fruiting pedicels | pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched. |
descending to horizontal, usually recurved, rarely straight, 4–20 mm, pubescent, trichomes appressed, 3–7-rayed. |
Fruits | closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm; valves pubescent throughout; ovules 38–64 per ovary; style 0.05–0.1 mm. |
widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (6–)8–12 cm × 1.5–2.5 mm; valves glabrous or trichomes sometimes scattered; ovules 140–180 per ovary; style 0.05–0.3 mm. |
Seeds | uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.3 mm wide. |
uniseriate or sub-biseriate, 1.7–2 × 1.2–1.7 mm; wing continuous, 0.2–0.4 mm wide. |
2n | = 14. |
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Boechera puberula |
Boechera californica |
|
Phenology | Flowering Apr–Jul. | Flowering Mar–Jun. |
Habitat | Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities | Rocky slopes and gravelly soil in desert chaparral and oak woodlands |
Elevation | 1300-2900 m (4300-9500 ft) | 300-2300 m (1000-7500 ft) |
Distribution |
CA; NV; OR; UT
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CA; Mexico (Baja California)
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Discussion | Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera californica is an apomictic species that arose through hybridization between B. arcuata and B. pulchra (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). In the flora area, B. californica is known only from Los Angeles, Riverside, San Bernardino, and San Diego counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 399. | FNA vol. 7, p. 369. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis puberula, Arabis arida, Arabis beckwithii, Arabis lignipes var. impar, Arabis sabulosa, Arabis subpinnatifida var. beckwithii, Arabis subpinnatifida var. impar, B. beckwithii | Arabis sparsiflora var. californica |
Name authority | (Nuttall) Dorn: Brittonia, 55: 3. (2003) | (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 260. (2007) |
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