Boechera perennans |
Boechera williamsii |
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perennial rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody. | Perennials; short-lived; sexual; caudex not woody. |
Stems | usually 2–5 per caudex branch, arising laterally proximal to sterile shoots or rosette, often elevated above ground surface on woody base, (1.5–)2–7 dm, densely pubescent proximally, trichomes short-stalked, Y-shaped, 0.2–0.4 mm, often mixed with 3–5-rayed or (rarely) simple ones, usually glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
Basal leaves | blade oblanceolate to obovate, 3–15(–20) mm wide, margins dentate, ciliate proximally, trichomes to 1.2 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 3–6-rayed, 0.2–0.4 mm. |
blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
Cauline leaves | 4–12(–17), not concealing stem; blade auricles 0.5–3.5 mm, surfaces of distalmost leaves glabrous. |
6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 16–35-flowered, usually unbranched. |
5–17-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals white to purplish, 5–9 × 1–1.5 mm, glabrous or sparsely pubescent (occasionally some trichomes abaxially); pollen ellipsoid. |
erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually horizontal, straight or slightly recurved, (6–)10–25 mm, usually glabrous. |
erect, straight, 2–4(–7) mm, glabrous. |
Fruits | widely pendent, not appressed to rachis, not secund, usually curved, edges parallel, (3–)4–7 cm × 1.7–2.1 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.4 mm. |
erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 1.1–1.5 × 0.9–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
2n | = 14. |
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Boechera perennans |
Boechera williamsii |
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Phenology | Flowering Feb–May. | Flowering May–Jun. |
Habitat | Rocky slopes and gravelly soil in warm desert, chaparral, low montane habitats | Gravelly soil in sagebrush-grassland communities |
Elevation | 200-1700 m (700-5600 ft) | 2300-2800 m (7500-9200 ft) |
Distribution |
AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Sonora)
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WY |
Discussion | The circumscription of Boechera perennans followed here is much narrower than that adopted by R. C. Rollins (1993). The most significant difference is the removal of Arabis gracilenta. Recent studies (M. D. Windham and Allphin, in prep.) indicate that the latter represents an older name for the species usually called A. (Boechera) selbyi. Although superficially similar, B. perennans is distinguished from B. gracilenta by having stems arising from above ground surface on woody bases, proximal stems with mostly Y-shaped trichomes, strongly dentate margins on basal leaves, and uniseriate seeds. Typically, B. perennans is a sexual diploid largely restricted to the warm deserts of Arizona, California, Nevada, and southern New Mexico. Plants of B. gracilenta are scattered across the Colorado Plateau (northeast of the range of B. perennans) and are apomictic triploids, apparently produced by hybridization between B. fendleri and B. pallidifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 396. | FNA vol. 7, p. 411. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis perennans, Arabis angulata, Arabis arcuata var. perennans, Arabis eremophila, Arabis recondita | Arabis williamsii |
Name authority | (S. Watson) W. A. Weber: Phytologia 51: 370. (1982) | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) |
Web links |