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perennial rockcress

Shevock's rockcress

Habit Perennials; long-lived; sexual; caudex woody. Perennials; long-lived; (pulvinate); sexual; caudex woody (with persistent, crowded leaf bases).
Stems

usually 2–5 per caudex branch, arising laterally proximal to sterile shoots or rosette, often elevated above ground surface on woody base, (1.5–)2–7 dm, densely pubescent proximally, trichomes short-stalked, Y-shaped, 0.2–0.4 mm, often mixed with 3–5-rayed or (rarely) simple ones, usually glabrous distally.

1 per caudex branch, arising from center of rosette near ground surface, 0.5–1 dm, puberulent proximally, trichomes (scattered), simple, 0.02–0.1 mm, similarly pubescent distally.

Basal leaves

blade oblanceolate to obovate, 3–15(–20) mm wide, margins dentate, ciliate proximally, trichomes to 1.2 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 3–6-rayed, 0.2–0.4 mm.

blade oblong-oblanceolate, 1–2 mm wide, margins entire, ciliate distally, trichomes (simple), 0.02–0.1 mm, surfaces glabrous.

Cauline leaves

4–12(–17), not concealing stem;

blade auricles 0.5–3.5 mm, surfaces of distalmost leaves glabrous.

3–7, not concealing stem;

blade auricles absent, surfaces of distalmost leaves glabrous.

Racemes

16–35-flowered, usually unbranched.

4–7-flowered, unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals white to purplish, 5–9 × 1–1.5 mm, glabrous or sparsely pubescent (occasionally some trichomes abaxially);

pollen ellipsoid.

ascending at anthesis;

sepals glabrous;

petals lavender, 4–5 × 1–1.2 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

usually horizontal, straight or slightly recurved, (6–)10–25 mm, usually glabrous.

ascending, straight, 5–9 mm, glabrous.

Fruits

widely pendent, not appressed to rachis, not secund, usually curved, edges parallel, (3–)4–7 cm × 1.7–2.1 mm;

valves glabrous;

ovules 60–96 per ovary;

style 0.05–0.4 mm.

ascending, not appressed to rachis, not secund, straight, edges parallel, 2.5–3 cm × 1–1.5 mm;

valves glabrous;

ovules 30–34 per ovary;

style ca. 0.5 mm.

Seeds

uniseriate, 1.1–1.5 × 0.9–1.2 mm;

wing continuous, 0.1–0.2 mm wide.

uniseriate, ca. 1.2 × 0.8 mm;

wing distal or continuous, ca. 0.1 mm wide.

2n

= 14.

Boechera perennans

Boechera shevockii

Phenology Flowering Feb–May. Flowering Jun.
Habitat Rocky slopes and gravelly soil in warm desert, chaparral, low montane habitats Ledges of rock outcrops
Elevation 200-1700 m (700-5600 ft) ca. 2500 m (ca. 8200 ft)
Distribution
from FNA
AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

The circumscription of Boechera perennans followed here is much narrower than that adopted by R. C. Rollins (1993). The most significant difference is the removal of Arabis gracilenta. Recent studies (M. D. Windham and Allphin, in prep.) indicate that the latter represents an older name for the species usually called A. (Boechera) selbyi. Although superficially similar, B. perennans is distinguished from B. gracilenta by having stems arising from above ground surface on woody bases, proximal stems with mostly Y-shaped trichomes, strongly dentate margins on basal leaves, and uniseriate seeds. Typically, B. perennans is a sexual diploid largely restricted to the warm deserts of Arizona, California, Nevada, and southern New Mexico. Plants of B. gracilenta are scattered across the Colorado Plateau (northeast of the range of B. perennans) and are apomictic triploids, apparently produced by hybridization between B. fendleri and B. pallidifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera shevockii is a close relative of B. davidsonii, from which it is readily distinguished by having puberulent (versus glabrous) stems, shorter (4–5 versus 6–10 mm) petals, and much shorter (5–7 versus 30–80 mm) basal leaves. It is known only from the type specimen, from the southern Sierra Nevada in Tulare County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 396. FNA vol. 7, p. 406.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis perennans, Arabis angulata, Arabis arcuata var. perennans, Arabis eremophila, Arabis recondita
Name authority (S. Watson) W. A. Weber: Phytologia 51: 370. (1982) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 82. (2006)
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