Boechera perennans |
Boechera perstellata |
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perennial rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody. | Perennials; usually short-lived; sexual; caudex not woody. |
Stems | usually 2–5 per caudex branch, arising laterally proximal to sterile shoots or rosette, often elevated above ground surface on woody base, (1.5–)2–7 dm, densely pubescent proximally, trichomes short-stalked, Y-shaped, 0.2–0.4 mm, often mixed with 3–5-rayed or (rarely) simple ones, usually glabrous distally. |
usually 2–5 per plant, arising from margin of rosette near ground surface, (1–)2–8 dm, densely pubescent proximally, trichomes short-stalked, 3- or 4-rayed, 0.1–0.5 mm, similarly pubescent distally. |
Basal leaves | blade oblanceolate to obovate, 3–15(–20) mm wide, margins dentate, ciliate proximally, trichomes to 1.2 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 3–6-rayed, 0.2–0.4 mm. |
blade obovate to oblanceolate, (5–)10–45 mm wide, margins usually coarsely dentate, rarely sinuate, ciliate on petiole base, surfaces moderately pubescent, trichomes short-stalked, 3- or 4(–6)-rayed, 0.1–0.5 mm. |
Cauline leaves | 4–12(–17), not concealing stem; blade auricles 0.5–3.5 mm, surfaces of distalmost leaves glabrous. |
6–25, concealing stem proximally; blade auricles 1–6 mm, surfaces of distalmost leaves pubescent. |
Racemes | 16–35-flowered, usually unbranched. |
10–40-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals white to purplish, 5–9 × 1–1.5 mm, glabrous or sparsely pubescent (occasionally some trichomes abaxially); pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals purplish, 3–4 × 0.7–1 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually horizontal, straight or slightly recurved, (6–)10–25 mm, usually glabrous. |
usually horizontal, usually straight, rarely slightly curved, 5–15 mm, pubescent, trichomes spreading, simple. |
Fruits | widely pendent, not appressed to rachis, not secund, usually curved, edges parallel, (3–)4–7 cm × 1.7–2.1 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.4 mm. |
usually horizontal, not appressed to rachis, not secund, straight, edges parallel, (1.5–)2–3.3 cm × 0.8–1 mm; valves sparsely pubescent throughout; ovules 12–20 per ovary; style 0.4–0.8 mm. |
Seeds | uniseriate, 1.1–1.5 × 0.9–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
uniseriate, 0.8–1.1 × 0.6–0.8 mm, not winged. |
2n | = 14. |
= 14. |
Boechera perennans |
Boechera perstellata |
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Phenology | Flowering Feb–May. | Flowering Apr–May. |
Habitat | Rocky slopes and gravelly soil in warm desert, chaparral, low montane habitats | Calcareous bluffs and wooded hillsides |
Elevation | 200-1700 m (700-5600 ft) | 200-400 m (700-1300 ft) |
Distribution |
AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Sonora)
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KY; TN |
Discussion | The circumscription of Boechera perennans followed here is much narrower than that adopted by R. C. Rollins (1993). The most significant difference is the removal of Arabis gracilenta. Recent studies (M. D. Windham and Allphin, in prep.) indicate that the latter represents an older name for the species usually called A. (Boechera) selbyi. Although superficially similar, B. perennans is distinguished from B. gracilenta by having stems arising from above ground surface on woody bases, proximal stems with mostly Y-shaped trichomes, strongly dentate margins on basal leaves, and uniseriate seeds. Typically, B. perennans is a sexual diploid largely restricted to the warm deserts of Arizona, California, Nevada, and southern New Mexico. Plants of B. gracilenta are scattered across the Colorado Plateau (northeast of the range of B. perennans) and are apomictic triploids, apparently produced by hybridization between B. fendleri and B. pallidifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Boechera perstellata is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 396. | FNA vol. 7, p. 396. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis perennans, Arabis angulata, Arabis arcuata var. perennans, Arabis eremophila, Arabis recondita | Arabis perstellata, Arabis perstellata var. ampla |
Name authority | (S. Watson) W. A. Weber: Phytologia 51: 370. (1982) | (E. L. Braun) Al-Shehbaz: Novon 13: 388. (2003) |
Web links |