Boechera perennans |
Boechera lyallii |
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perennial rockcress |
jewelled suncress, Lyall's rockcress, Lyall's slender rock-cress, murry's rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody. | Perennials; long-lived; (cespitose); sexual or apomictic; caudex woody. |
Stems | usually 2–5 per caudex branch, arising laterally proximal to sterile shoots or rosette, often elevated above ground surface on woody base, (1.5–)2–7 dm, densely pubescent proximally, trichomes short-stalked, Y-shaped, 0.2–0.4 mm, often mixed with 3–5-rayed or (rarely) simple ones, usually glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.3–1.5(–2) dm, glabrous throughout. |
Basal leaves | blade oblanceolate to obovate, 3–15(–20) mm wide, margins dentate, ciliate proximally, trichomes to 1.2 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 3–6-rayed, 0.2–0.4 mm. |
blade linear-oblanceolate, 1–5(–8) mm wide, margins entire, usually ciliate at least proximally, trichomes (simple and/or short-stalked, 2- or 3-rayed), 0.1–0.3 mm, surfaces (sometimes the entire leaf) usually glabrous or (rarely, the youngest leaves of sterile shoots) pubescent, trichomes 4–6-rayed, 0.05–0.1 mm. |
Cauline leaves | 4–12(–17), not concealing stem; blade auricles 0.5–3.5 mm, surfaces of distalmost leaves glabrous. |
1–5, usually not concealing stem; blade auricles 0.5–1.5 mm or, rarely, absent, surfaces glabrous. |
Racemes | 16–35-flowered, usually unbranched. |
2–10(–15)-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals white to purplish, 5–9 × 1–1.5 mm, glabrous or sparsely pubescent (occasionally some trichomes abaxially); pollen ellipsoid. |
erect at anthesis; sepals glabrous; petals lavender to purplish, 6–8.5 × 1.5–3 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | usually horizontal, straight or slightly recurved, (6–)10–25 mm, usually glabrous. |
erect, straight, 3–8(–15) mm, glabrous. |
Fruits | widely pendent, not appressed to rachis, not secund, usually curved, edges parallel, (3–)4–7 cm × 1.7–2.1 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.4 mm. |
erect, appressed to rachis, not secund, straight, edges parallel, 3–5.6 cm × 1.5–2.5 mm; valves glabrous; ovules 34–64 per ovary; style 0.1–0.7 mm. |
Seeds | uniseriate, 1.1–1.5 × 0.9–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
biseriate or sub-biseriate, 1.5–2 × 1–1.5 mm; wing continuous, 0.3–0.5 mm wide. |
2n | = 14. |
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Boechera perennans |
Boechera lyallii |
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Phenology | Flowering Feb–May. | Flowering Jun–Aug. |
Habitat | Rocky slopes and gravelly soil in warm desert, chaparral, low montane habitats | Cliffs, talus slopes, gravelly soil in alpine and subalpine habitats |
Elevation | 200-1700 m (700-5600 ft) | 1400-3700 m (4600-12100 ft) |
Distribution |
AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Sonora)
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CA; ID; MT; NV; OR; UT; WA; BC; YT
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Discussion | The circumscription of Boechera perennans followed here is much narrower than that adopted by R. C. Rollins (1993). The most significant difference is the removal of Arabis gracilenta. Recent studies (M. D. Windham and Allphin, in prep.) indicate that the latter represents an older name for the species usually called A. (Boechera) selbyi. Although superficially similar, B. perennans is distinguished from B. gracilenta by having stems arising from above ground surface on woody bases, proximal stems with mostly Y-shaped trichomes, strongly dentate margins on basal leaves, and uniseriate seeds. Typically, B. perennans is a sexual diploid largely restricted to the warm deserts of Arizona, California, Nevada, and southern New Mexico. Plants of B. gracilenta are scattered across the Colorado Plateau (northeast of the range of B. perennans) and are apomictic triploids, apparently produced by hybridization between B. fendleri and B. pallidifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The taxon traditionally treated as Arabis (Boechera) lyallii var. nubigena is here segregated as B. paupercula (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). Completely glabrous individuals of B. lyallii are sometimes confused with B. davidsonii, but they are easily distinguished by the absence of persistent leaf bases on caudex branches, erect and appressed (versus ascending) fruits, and biseriate to sub-biseriate (versus uniseriate) seeds. Both sexual and apomictic collections are known; further study is needed to determine whether they truly are conspecific. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 396. | FNA vol. 7, p. 389. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis perennans, Arabis angulata, Arabis arcuata var. perennans, Arabis eremophila, Arabis recondita | Arabis lyallii, Arabis armerifolia, Arabis densa, Arabis drummondii var. alpina, Arabis drummondii var. lyallii, Arabis multiceps, Arabis murrayi |
Name authority | (S. Watson) W. A. Weber: Phytologia 51: 370. (1982) | (S. Watson) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) |
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