Boechera peirsonii |
Boechera divaricarpa |
|
---|---|---|
Peirson's rockcress, San Bernardino Rock cress |
spreading-pod rock-cress, uplifting suncress |
|
Habit | Perennials; long-lived; (cespitose); apomictic; caudex woody. | Biennials or perennials; short-lived; apomictic; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–2.5 dm, glabrous or sparsely pubescent proximally, trichomes short-stalked, 2–5-rayed (rarely, some simple), 0.4–0.6 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)3–9 dm, glabrous or pubescent proximally, trichomes sessile, 2–4-rayed (some simple), to 0.7 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 2.5–6 mm wide, margins entire, ciliate proximally, trichomes (simple), surfaces densely pubescent, trichomes short-stalked, 4–7-rayed, 0.3–0.5 mm. |
blade oblanceolate, 2–10 mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes sessile, 2–6-rayed, 0.1–0.4 mm. |
Cauline leaves | 3–12, often concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves usually glabrous, sometimes margins ciliate. |
(10–)15–56, concealing stem proximally; blade auricles 1–5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 12–25-flowered, usually unbranched. |
12–40(–65)-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 5–6 × 1.5–2 mm, glabrous; pollen spheroid. |
divaricate-ascending at anthesis; sepals glabrous or with scattered trichomes; petals usually purple, rarely lavender, 6–9 × 1.5–3 mm, glabrous; pollen spheroid. |
Fruiting pedicels | ascending to divaricate-descending, straight, 2–6 mm, glabrous. |
divaricate-ascending to horizontal, straight, 5–12 mm, glabrous. |
Fruits | ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 2–3.7 cm × 2–2.8 mm; valves glabrous; ovules 56–80 per ovary; style 0.1–0.3 mm. |
divaricate-ascending to horizontal, not appressed to rachis, not secund, straight, edges parallel, (4.5–)5.5–11 cm × 1.7–2.5 mm; valves glabrous; ovules 114–142 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 1–1.5 × 0.8–1 mm; wing distal or continuous, 0.05–0.1 mm wide. |
uniseriate to sub-biseriate, 1.4–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 21. |
|
Boechera peirsonii |
Boechera divaricarpa |
|
Phenology | Flowering Jun–Sep. | Flowering May–Jul. |
Habitat | Granitic ledges and talus slopes | Rock outcrops, talus slopes and gravelly hillsides in sagebrush, mountain shrub, and open conifer forests |
Elevation | 2700-3400 m (8900-11200 ft) | 900-2500 m (3000-8200 ft) |
Distribution |
CA |
CA; ID; MT; NV; UT; WA; WY
|
Discussion | Boechera peirsonii is an apomictic species of unknown origin; although previously treated as a variety of Arabis (B.) breweri, it is quite distinct from that taxon (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). All known collections come from the San Bernardino Mountains of southern California, ca. 200 km distant from the nearest populations of B. breweri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The name Arabis (Boechera) divaricarpa has been applied to nearly every hybrid containing a genome derived from B. stricta. This presents a serious barrier to understanding the evolution of Boechera and also is contrary to the International Code of Botanical Nomenclature, because some names usually placed in synonymy (i.e., B. grahamii and B. brachycarpa) have priority at species level (M. D. Windham and I. A. Al-Shehbaz 2007b). To address this problem, we treat the following as distinct species: B. acutina, B. grahamii (= B. brachycarpa of R. D. Dorn 2001), and B. pratincola (all considered synonyms of A. divaricarpa by R. C. Rollins 1993), and B. calderi, B. elkoensis, and B. quebecensis (taxa described after 1993). Detailed comparison among these taxa are provided by Windham and Al-Shehbaz (2007, 2007b). The narrow concept of B. divaricarpa advocated here encompasses apomictic triploid populations containing three distinct genomes, one each derived from B. retrofracta, B. sparsiflora, and B. stricta. If the species is defined more broadly, the name B. grahamii has priority. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 394. | FNA vol. 7, p. 374. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis breweri var. pecuniaria | Arabis divaricarpa, Arabis stokesiae |
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 270. (2007) | (A. Nelson) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) |
Web links |
|
|