Boechera paupercula |
Boechera pinetorum |
|
---|---|---|
boechera paupercula, littleleaf rockcress, small-flower rock-cress, tiny suncress |
Holboell's rock-cress, woodland rock-cress |
|
Habit | Perennials; long-lived; (cespitose); sexual; caudex often woody. | Perennials; short-lived; apomictic; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.3–1.5 dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, (0.07–)0.1–0.2 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 2–8(–9.6) dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
Basal leaves | blade linear-oblanceolate, 1–3(–5) mm wide, margins entire, ciliate proximally, trichomes to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 2–6-rayed, (0.07–)0.1–0.2 mm. |
blade oblanceolate, 2–8(–11) mm wide, margins entire or denticulate, ciliate proximally, trichomes (simple), 0.5–1 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed, 0.2–0.5 mm, these mixed with fewer simple ones. |
Cauline leaves | 2–6, rarely concealing stem; blade auricles 0 (or 0.2–1.5) mm, surfaces of distalmost leaves glabrous. |
7–33, often concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves sparsely pubescent or glabrous. |
Racemes | 3–8-flowered, unbranched. |
15–63-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous or sparsely pubescent; petals lavender to purplish, 4–6(–7) × 1–2 mm, glabrous; pollen ellipsoid. |
divaricate to pendent at anthesis; sepals pubescent; petals white to lavender, 5–6 × 1–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect, straight, 3–9 mm, glabrous or sparsely pubescent. |
reflexed, abruptly recurved near base, otherwise straight, 4–12 mm, sparsely pubescent or glabrous, trichomes spreading, simple and 2-rayed. |
Fruits | erect, appressed to rachis, not secund, straight, edges parallel, 2.5–5.5 cm × 1.3–1.7(–2) mm; valves glabrous; ovules 24–40 per ovary; style 0.2–1 mm. |
reflexed to closely pendent, not appressed to rachis, not secund, straight or curved, edges parallel, (4.5–)5.5–8.5 cm × 1.5–2 mm; valves glabrous; ovules 70–110 per ovary; style 0.05–0.4 mm. |
Seeds | usually uniseriate, rarely sub-biseriate, 1.5–2(–2.5) × 1–1.4 mm; wing continuous, 0.3–1 mm wide. |
uniseriate, 1.5–1.8 × 1.2–1.5 mm; wing continuous, 0.15–0.3 mm wide. |
Boechera paupercula |
Boechera pinetorum |
|
Phenology | Flowering Jun–Aug. | Flowering May–Jul. |
Habitat | Rock outcrops, talus slopes, gravelly soil in alpine and subalpine habitats | Rock outcrops and gravelly soil in meadows and open conifer forests |
Elevation | 2400-3400 m (7900-11200 ft) | 1100-3200 m (3600-10500 ft) |
Distribution |
CA; ID; OR; WA; WY
|
CA; NV; OR
|
Discussion | Boechera paupercula is usually subsumed under Arabis (Boechera) lyallii but is amply distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most authors (e.g., R. C. Rollins 1993; R. D. Dorn 2001; S. L. Welsh et al. 2003; N. H. Holmgren 2005b) have treated Boechera pinetorum as a variety of Arabis (Boechera) holboellii. Under this guise, the name has been applied to a vast array of plants collected throughout western North America. This includes a diversity of sexual diploids plus nearly every hybrid containing a genome from B. retrofracta. Based on re-examination of the type collection, we have adopted a much narrower concept of the species. Morphological evidence suggests that B. pinetorum is an apomictic triploid hybrid containing three different genomes, derived from B. rectissima, B. retrofracta, and B. sparsiflora. Plants closely resembling the type of B. pinetorum are confined to the northern Sierra Nevada and adjacent southern Cascades. The majority of collections previously associated with the epithet pinetorum represent B. pauciflora (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 394. | FNA vol. 7, p. 397. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis paupercula, Arabis lyallii var. nubigena, Arabis microphylla var. nubigena, Arabis nubigena | Arabis pinetorum, Arabis divaricarpa var. pinetorum, Arabis holboellii var. pinetorum, B. holboellii var. pinetorum |
Name authority | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 75. (2006) | (Tidestrom) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 271. (2007) |
Web links |
|
|