Boechera paupercula |
Boechera perennans |
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boechera paupercula, littleleaf rockcress, small-flower rock-cress, tiny suncress |
perennial rockcress |
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Habit | Perennials; long-lived; (cespitose); sexual; caudex often woody. | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.3–1.5 dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, (0.07–)0.1–0.2 mm, glabrous or sparsely pubescent distally. |
usually 2–5 per caudex branch, arising laterally proximal to sterile shoots or rosette, often elevated above ground surface on woody base, (1.5–)2–7 dm, densely pubescent proximally, trichomes short-stalked, Y-shaped, 0.2–0.4 mm, often mixed with 3–5-rayed or (rarely) simple ones, usually glabrous distally. |
Basal leaves | blade linear-oblanceolate, 1–3(–5) mm wide, margins entire, ciliate proximally, trichomes to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 2–6-rayed, (0.07–)0.1–0.2 mm. |
blade oblanceolate to obovate, 3–15(–20) mm wide, margins dentate, ciliate proximally, trichomes to 1.2 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 3–6-rayed, 0.2–0.4 mm. |
Cauline leaves | 2–6, rarely concealing stem; blade auricles 0 (or 0.2–1.5) mm, surfaces of distalmost leaves glabrous. |
4–12(–17), not concealing stem; blade auricles 0.5–3.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 3–8-flowered, unbranched. |
16–35-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous or sparsely pubescent; petals lavender to purplish, 4–6(–7) × 1–2 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white to purplish, 5–9 × 1–1.5 mm, glabrous or sparsely pubescent (occasionally some trichomes abaxially); pollen ellipsoid. |
Fruiting pedicels | erect, straight, 3–9 mm, glabrous or sparsely pubescent. |
usually horizontal, straight or slightly recurved, (6–)10–25 mm, usually glabrous. |
Fruits | erect, appressed to rachis, not secund, straight, edges parallel, 2.5–5.5 cm × 1.3–1.7(–2) mm; valves glabrous; ovules 24–40 per ovary; style 0.2–1 mm. |
widely pendent, not appressed to rachis, not secund, usually curved, edges parallel, (3–)4–7 cm × 1.7–2.1 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.4 mm. |
Seeds | usually uniseriate, rarely sub-biseriate, 1.5–2(–2.5) × 1–1.4 mm; wing continuous, 0.3–1 mm wide. |
uniseriate, 1.1–1.5 × 0.9–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 14. |
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Boechera paupercula |
Boechera perennans |
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Phenology | Flowering Jun–Aug. | Flowering Feb–May. |
Habitat | Rock outcrops, talus slopes, gravelly soil in alpine and subalpine habitats | Rocky slopes and gravelly soil in warm desert, chaparral, low montane habitats |
Elevation | 2400-3400 m (7900-11200 ft) | 200-1700 m (700-5600 ft) |
Distribution |
CA; ID; OR; WA; WY
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AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Sonora)
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Discussion | Boechera paupercula is usually subsumed under Arabis (Boechera) lyallii but is amply distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The circumscription of Boechera perennans followed here is much narrower than that adopted by R. C. Rollins (1993). The most significant difference is the removal of Arabis gracilenta. Recent studies (M. D. Windham and Allphin, in prep.) indicate that the latter represents an older name for the species usually called A. (Boechera) selbyi. Although superficially similar, B. perennans is distinguished from B. gracilenta by having stems arising from above ground surface on woody bases, proximal stems with mostly Y-shaped trichomes, strongly dentate margins on basal leaves, and uniseriate seeds. Typically, B. perennans is a sexual diploid largely restricted to the warm deserts of Arizona, California, Nevada, and southern New Mexico. Plants of B. gracilenta are scattered across the Colorado Plateau (northeast of the range of B. perennans) and are apomictic triploids, apparently produced by hybridization between B. fendleri and B. pallidifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 394. | FNA vol. 7, p. 396. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis paupercula, Arabis lyallii var. nubigena, Arabis microphylla var. nubigena, Arabis nubigena | Arabis perennans, Arabis angulata, Arabis arcuata var. perennans, Arabis eremophila, Arabis recondita |
Name authority | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 75. (2006) | (S. Watson) W. A. Weber: Phytologia 51: 370. (1982) |
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