Boechera paupercula |
Boechera pendulocarpa |
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boechera paupercula, littleleaf rockcress, small-flower rock-cress, tiny suncress |
danglepod rockcress, drop-seed rock-cress, earless suncress, Holboell's rockcress |
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Habit | Perennials; long-lived; (cespitose); sexual; caudex often woody. | Perennials; short- to long-lived; sexual; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.3–1.5 dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, (0.07–)0.1–0.2 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–3 dm, densely pubescent proximally, trichomes simple and stalked, 2–4-rayed, 0.1–0.4 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade linear-oblanceolate, 1–3(–5) mm wide, margins entire, ciliate proximally, trichomes to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 2–6-rayed, (0.07–)0.1–0.2 mm. |
blade narrowly oblanceolate, 1.5–4(–5) mm wide, margins entire, often ciliate along petiole, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.08–0.2 mm. |
Cauline leaves | 2–6, rarely concealing stem; blade auricles 0 (or 0.2–1.5) mm, surfaces of distalmost leaves glabrous. |
6–17, sometimes concealing stem proximally; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 3–8-flowered, unbranched. |
4–11-flowered, unbranched. |
Flowers | erect at anthesis; sepals glabrous or sparsely pubescent; petals lavender to purplish, 4–6(–7) × 1–2 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white or lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | erect, straight, 3–9 mm, glabrous or sparsely pubescent. |
arched, gently recurved, 3–8(–12) mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | erect, appressed to rachis, not secund, straight, edges parallel, 2.5–5.5 cm × 1.3–1.7(–2) mm; valves glabrous; ovules 24–40 per ovary; style 0.2–1 mm. |
pendent, not appressed to rachis, not secund, usually straight, edges parallel, (2–)2.5–3.8 cm × 1.5–2.2 mm; valves glabrous; ovules 66–92 per ovary; style 0.3–0.5 mm. |
Seeds | usually uniseriate, rarely sub-biseriate, 1.5–2(–2.5) × 1–1.4 mm; wing continuous, 0.3–1 mm wide. |
sub-biseriate, 1–1.2 × 0.7–0.9 mm; wing distal, 0.05–0.1 mm wide. |
2n | = 14. |
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Boechera paupercula |
Boechera pendulocarpa |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Jul. |
Habitat | Rock outcrops, talus slopes, gravelly soil in alpine and subalpine habitats | Rock outcrops and gravelly slopes in sagebrush, open conifer forests, subalpine meadows |
Elevation | 2400-3400 m (7900-11200 ft) | 1000-3300 m (3300-10800 ft) |
Distribution |
CA; ID; OR; WA; WY
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CA; ID; MT; NV; OR; UT; WY; BC; YT
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Discussion | Boechera paupercula is usually subsumed under Arabis (Boechera) lyallii but is amply distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993), B. pendulocarpa is easily distinguished from that species by having simple and 2–4-rayed (versus 4–8-rayed) trichomes proximally on stems, cauline leaves without auricles, fruiting pedicels gently (versus sharply) recurved, and shorter (2–)2.5–3.8 (versus 3.5–6.5) cm, non-secund fruits. The two taxa have allopatric distributions, with B. pendulocarpa found in the mountains of western North America and B. holboellii apparently confined to Greenland. Recent use of the name A. (Boechera) exilis for this taxon (e.g., G. A. Mulligan 1996; R. D. Dorn 2001; N. H. Holmgren 2005b) is based on misinterpretation of the type (M. D. Windham and I. A. Al-Shehbaz 2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 394. | FNA vol. 7, p. 395. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis paupercula, Arabis lyallii var. nubigena, Arabis microphylla var. nubigena, Arabis nubigena | Arabis pendulocarpa, Arabis holboellii var. pendulocarpa, B. holboellii var. pendulocarpa |
Name authority | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 75. (2006) | (A. Nelson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 77. (2006) |
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