Boechera paupercula |
Boechera inyoensis |
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boechera paupercula, littleleaf rockcress, small-flower rock-cress, tiny suncress |
Inyo rockcress |
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Habit | Perennials; long-lived; (cespitose); sexual; caudex often woody. | Perennials; short-lived; apomictic; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.3–1.5 dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, (0.07–)0.1–0.2 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex, arising from center of rosette near ground surface, (1–)2.5–6.5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
Basal leaves | blade linear-oblanceolate, 1–3(–5) mm wide, margins entire, ciliate proximally, trichomes to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 2–6-rayed, (0.07–)0.1–0.2 mm. |
blade oblanceolate, 1–4(–8) mm wide, margins entire, ciliate along petiole and proximally, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 3–10-rayed, 0.2–0.7 mm. |
Cauline leaves | 2–6, rarely concealing stem; blade auricles 0 (or 0.2–1.5) mm, surfaces of distalmost leaves glabrous. |
(7–)12–35, concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves densely pubescent. |
Racemes | 3–8-flowered, unbranched. |
10–65-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous or sparsely pubescent; petals lavender to purplish, 4–6(–7) × 1–2 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender to purplish, 5–8 × 1.2–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect, straight, 3–9 mm, glabrous or sparsely pubescent. |
divaricate-ascending to horizontal, straight or slightly curved downward, 5–15 mm, sparsely pubescent, trichomes appressed, branched. |
Fruits | erect, appressed to rachis, not secund, straight, edges parallel, 2.5–5.5 cm × 1.3–1.7(–2) mm; valves glabrous; ovules 24–40 per ovary; style 0.2–1 mm. |
divaricate-ascending to widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.7–6.5 cm × 1.5–2.2 mm; valves glabrous; ovules 74–134 per ovary; style 0.05–0.2 mm. |
Seeds | usually uniseriate, rarely sub-biseriate, 1.5–2(–2.5) × 1–1.4 mm; wing continuous, 0.3–1 mm wide. |
usually sub-biseriate, rarely uniseriate, 1.7–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 21. |
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Boechera paupercula |
Boechera inyoensis |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Jun. |
Habitat | Rock outcrops, talus slopes, gravelly soil in alpine and subalpine habitats | Limestone and volcanic rock outcrops and clay soils in desert scrub and pinyon-juniper woodlands |
Elevation | 2400-3400 m (7900-11200 ft) | 1400-2400 m (4600-7900 ft) |
Distribution |
CA; ID; OR; WA; WY
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CA; NV; UT
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Discussion | Boechera paupercula is usually subsumed under Arabis (Boechera) lyallii but is amply distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Some plants here assigned to Boechera inyoensis were treated by R. C. Rollins (1993) and N. H. Holmgren (2005b) as Arabis (or B.) beckwithii. Closer examination of the type specimens of A. beckwithii reveals that they are conspecific with B. puberula, a sexual diploid widespread in the Great Basin of western North America. Boechera inyoensis is an apomictic triploid that clearly contains at least one genome derived from B. shockleyi (M. D. Windham and I. A. Al-Shehbaz 2007b); the other parent has not been determined. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 394. | FNA vol. 7, p. 385. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis paupercula, Arabis lyallii var. nubigena, Arabis microphylla var. nubigena, Arabis nubigena | Arabis inyoensis, Arabis holboellii var. derensis, B. selbyi var. inyoensis |
Name authority | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 75. (2006) | (Rollins) Al-Shehbaz: Novon 13: 386. (2003) |
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