Boechera pauciflora |
Boechera fructicosa |
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Columbia rockcress, few-flower rock-cress, fuzzy suncress, hairy-stem rock-cress, small-flower rockcress |
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Habit | Perennials; short- to long-lived; apomictic; caudex sometimes woody. | Perennials; short-lived; apomictic; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.4–)3–11.2 dm, densely pubescent proximally, trichomes simple, 0.6–1.5 mm, mixed with stalked, 2- (or 3-)rayed ones, 0.2–0.4 mm, sparsely pubescent distally. |
3 to many per caudex branch, arising from margin of rosette near ground surface, 3.5–6 dm, pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.2–0.3 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 3–10 mm wide, margins usually dentate, rarely entire, sometimes ciliate, trichomes (simple or branched), to 1 mm, surfaces densely pubescent, trichomes stalked, 2–5-rayed, 0.3–0.6 mm. |
blade oblanceolate, 3–7 mm wide, margins dentate, ciliate proximally, trichomes (simple), to 0.3 mm, surfaces pubescent, trichomes short-stalked, 4–8-rayed, 0.05–0.2 mm. |
Cauline leaves | (8–)14–60, often concealing stem proximally; blade auricles (1–)3–10 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
5–10, not concealing stem; blade auricles 0.5–1 mm, surfaces of distalmost leaves glabrous. |
Racemes | 17–60-flowered, usually unbranched. |
10–18-flowered, usually unbranched. |
Flowers | divaricate at anthesis; sepals pubescent; petals lavender to whitish, 5–8 × 1–2 mm, glabrous; pollen spheroid. |
divaricate-ascending at anthesis; sepals pubescent; petals lavender, 4–6 × 1–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | horizontal to divaricate-descending, usually straight, rarely slightly recurved, 4–13 mm, pubescent, trichomes spreading, 2- or 3-rayed. |
divaricate-descending, straight, 5–10 mm, glabrous. |
Fruits | horizontal, divaricate-descending or widely pendent, not appressed to rachis, not secund, curved, edges parallel, 5.5–10.5 cm × 1.5–2.2 mm; valves glabrous; ovules 80–162 per ovary; style 0.05–0.5 mm. |
divaricate-descending, not appressed to rachis, not secund, curved, edges parallel, 4–6 cm × 1.5–1.8 mm; valves glabrous; ovules 60–70 per ovary; style 0.1–0.4 mm. |
Seeds | uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.25 mm wide. |
uniseriate, 1.5–2 × 1–1.3 mm; wing distal, ca. 0.1 mm wide. |
2n | = 21. |
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Boechera pauciflora |
Boechera fructicosa |
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Phenology | Flowering May–Jun. | Flowering Jun–Jul. |
Habitat | Rocky soil in sagebrush areas, mountain shrub communities, edges of conifer forests | Dry, disturbed soil |
Elevation | 600-2500 m (2000-8200 ft) | ca. 2000 m (ca. 6600 ft) |
Distribution |
CA; ID; MT; NV; OR; UT; WA; WY; BC
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WY |
Discussion | Morphological evidence suggests that Boechera pauciflora is an apomictic species that arose through hybridization between B. retrofracta and B. sparsiflora. Specimens of B. pauciflora are commonly identified as Arabis holboellii var. pinetorum (= B. pinetorum), a superficially similar species restricted to the northern Sierra Nevada and southern Cascade Range (see M. D. Windham and I. A. 2007 for detailed comparison). Arabis elegans A. Nelson (1900), not Tineo & Lojacono (1886) is an illegitimate name, sometimes found in synonymy with Boechera pauciflora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera fructicosa is known only from the type collection (A. & E. Nelson 5681) taken near Undine Falls in Park County. Multiple plants are represented (comprising the holotype and four isotypes), though one specimen with the same number at RM is B. stricta. Although the exact parentage of this apomictic hybrid remains in doubt, it is virtually certain that B. microphylla contributed at least one genome. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 393. | FNA vol. 7, p. 379. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Sisymbrium pauciflorum, Arabis arcuata var. subvillosa, Arabis columbiana, Arabis perelegans, Arabis sparsiflora var. columbiana, Arabis sparsiflora var. subvillosa, B. sparsiflora var. subvillosa | Arabis fructicosa |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 268. (2007) | (A. Nelson) Al-Shehbaz: Novon 13: 385. (2003) |
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