Boechera paddoensis |
Boechera subpinnatifida |
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Mount Adams rock-cress, Mt. Adams rock-cress |
ashy rock-cress, Klamath rockcress |
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Habit | Perennials; long-lived; (cespitose); apomictic; caudex usually woody. | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). |
Stems | 1–3 per caudex branch, arising from center of rosette near ground surface, or arising laterally proximal to sterile shoots, 1–2.5 dm, glabrous throughout. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–3(–6) mm wide, margins usually entire, rarely apically few-toothed, not ciliate, surfaces sparsely pubescent, trichomes short-stalked, 4–8-rayed, 0.08–0.2 mm. |
blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
Cauline leaves | 4–8, not concealing stem; blade auricles 1–2 mm, surfaces of distalmost leaves glabrous. |
(10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
Racemes | 5–12(–18)-flowered, usually unbranched. |
8–30-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals glabrous; petals lavender to purple, 4–5.5 × 1–1.5 mm, glabrous; pollen spheroid. |
divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, straight, 2–5(–10) mm, glabrous. |
reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, secund (often weakly so), straight, edges parallel, 3–5.5 cm × 1.5–2 mm; valves glabrous; ovules 42–54 per ovary; style 0.05–0.1 mm. |
pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
Seeds | uniseriate, 1.1–1.5 × 0.9–1.3 mm; wing continuous, 0.1–0.3 mm wide. |
uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
2n | = 14. |
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Boechera paddoensis |
Boechera subpinnatifida |
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Phenology | Flowering Jul–Aug. | Flowering Mar–May. |
Habitat | Rocky ridges | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities |
Elevation | 1500-2100 m (4900-6900 ft) | 800-2400 m (2600-7900 ft) |
Distribution |
OR; WA
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CA; ID; NV; OR; UT
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Discussion | Morphological evidence suggests that Boechera paddoensis is an apomictic species that arose through hybridization between B. lemmonii and B. lyallii (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). Boechera paddoensis is known only from the mountains of central Washington and northeastern Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 392. | FNA vol. 7, p. 409. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis lemmonii var. paddoensis | Arabis subpinnatifida |
Name authority | (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 268. (2007) | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) |
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