Boechera oxylobula |
Boechera laevigata |
|
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|
smooth rockcress |
|
Habit | Perennials; short- to long-lived; (cespitose); sexual; caudex usually not woody. | Biennials; short-lived; sexual; caudex not evident. |
Stems | usually 3–7 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.4–2.5 dm, glabrous or pubescent proximally, trichomes simple and short-stalked, 2-rayed, 0.1–0.4 mm, glabrous distally. |
1 per plant, arising from center of rosette near ground surface, (1.5–)3–11 dm, glabrous throughout. |
Basal leaves | blade linear to linear-oblanceolate, 1–2.5 mm wide, margins usually entire, rarely denticulate, often ciliate, trichomes (simple), 0.3–0.7 mm, surfaces glabrous or sparsely pubescent, trichomes short-stalked, 2- or 3-rayed, 0.1–0.4 mm. |
blade obovate to oblanceolate, (4–)10–40 mm wide, margins serrate or dentate, ciliate-mucronate on teeth, trichomes often minute, surfaces glabrous or sparsely pubescent, trichomes simple, 0.1–0.6 mm. |
Cauline leaves | 3–12, not concealing stem; blade auricles absent, surfaces of distalmost leaves usually glabrous, rarely sparsely pubescent. |
7–15, often concealing stem proximally; blade auricles 3–12(–17) mm, surfaces of distalmost leaves glabrous. |
Racemes | 2–12-flowered, unbranched. |
16–45-flowered, sometimes branched. |
Flowers | ascending-divaricate at anthesis; sepals glabrous or pubescent; petals white to pale lavender, 4–5 × 1.5–2 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals glabrous; petals white, 3–5 × 1–1.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | divaricate-ascending to horizontal, slightly to strongly recurved, 3–8 mm, glabrous or sparsely pubescent, trichomes (isolated), simple. |
suberect to divaricate-ascending, straight to slightly curved, 5–23 mm, glabrous. |
Fruits | pendent, not appressed to rachis, not or, rarely, weakly secund, straight, edges parallel, 1.5–3.5 cm × 1.2–2 mm; valves glabrous; ovules 28–44 per ovary; style 0.1–0.4 mm. |
divaricate-ascending, not appressed to rachis, rarely somewhat secund, curved, edges parallel, (4–)6–11.7 cm × 1–2 mm; valves glabrous; ovules 50–80 per ovary; style 0.1–0.7(–1) mm. |
Seeds | uniseriate, 0.9–1.2 × 0.6–1 mm; wing often continuous, 0.07–0.1 mm wide. |
uniseriate, 1.2–2.2 × 0.8–1.4 mm; wing continuous, 0.1–0.3 mm wide distally. |
2n | = 14. |
|
Boechera oxylobula |
Boechera laevigata |
|
Phenology | Flowering May–Jul. | Flowering Mar–May. |
Habitat | Cliffs, rocky slopes, gravelly soil in sagebrush and open conifer forests | Rocky bluffs, cedar glades, wooded hillsides, floodplains |
Elevation | 2100-3600 m (6900-11800 ft) | 100-500 m (300-1600 ft) |
Distribution |
CO |
AL; AR; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; MI; MO; NC; NH; NJ; NY; OH; OK; PA; SC; TN; VA; VT; WI; WV; ON; QC |
Discussion | Most of the collections assigned here have been called Arabis (Boechera) demissa by other authors (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b). Because the holotype of A. demissa is identical to B. oxylobula in nearly every way (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison), we treat them as conspecific. Boechera oxylobula is restricted to Garfield, Gunnison, Hinsdale, Lake, Mineral, Park, and Saguache counties in central Colorado. The taxon traditionally treated as A. (Boechera) demissa var. languida is here recognized as an apomictic species of hybrid origin (see Windham and Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The taxon sometimes treated as Arabis laevigata var. missouriensis is here recognized as a separate species (Boechera missouriensis) based on its significantly longer petals (5–10 versus 3–5 mm) and distinctive, lyrate-pinnatifid basal leaves that persist well beyond anthesis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 391. | FNA vol. 7, p. 386. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis oxylobula, Arabis aprica, Arabis demissa, Arabis rugocarpa, B. demissa | Turritis laevigata, Arabis hastata, Arabis heterophylla, Arabis laevigata, Arabis lyrifolia |
Name authority | (Greene) W. A. Weber: Phytologia 51: 370. (1982) | (Muhlenberg ex Willdenow) Al-Shehbaz: Novon 13: 386. (2003) |
Web links |