Boechera oxylobula |
Boechera fernaldiana |
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Habit | Perennials; short- to long-lived; (cespitose); sexual; caudex usually not woody. | Perennials; long-lived; (cespitose); sexual; caudex somewhat woody (sometimes with persistent, crowded leaf bases). | ||||
Stems | usually 3–7 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.4–2.5 dm, glabrous or pubescent proximally, trichomes simple and short-stalked, 2-rayed, 0.1–0.4 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette, near ground surface or slightly elevated on woody base, 1–3.8 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.04–0.1 mm, sometimes mixed with stalked, 2-rayed ones, to 0.5 mm, usually sparsely pubescent distally. |
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Basal leaves | blade linear to linear-oblanceolate, 1–2.5 mm wide, margins usually entire, rarely denticulate, often ciliate, trichomes (simple), 0.3–0.7 mm, surfaces glabrous or sparsely pubescent, trichomes short-stalked, 2- or 3-rayed, 0.1–0.4 mm. |
blade narrowly oblanceolate, 1–4 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.04–0.1 mm. |
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Cauline leaves | 3–12, not concealing stem; blade auricles absent, surfaces of distalmost leaves usually glabrous, rarely sparsely pubescent. |
5–10, not concealing stem; blade auricles 0.3–2 mm, surfaces of distalmost leaves usually sparsely pubescent. |
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Racemes | 2–12-flowered, unbranched. |
10–20-flowered, usually unbranched. |
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Flowers | ascending-divaricate at anthesis; sepals glabrous or pubescent; petals white to pale lavender, 4–5 × 1.5–2 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals (purplish or greenish) glabrous or sparsely or moderately pubescent; petals white or purple to lavender, 7–12 × 1.5–4 mm, glabrous; pollen ellipsoid. |
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Fruiting pedicels | divaricate-ascending to horizontal, slightly to strongly recurved, 3–8 mm, glabrous or sparsely pubescent, trichomes (isolated), simple. |
divaricate-ascending, straight, 5–20 mm, sparsely pubescent, trichomes appressed, branched. |
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Fruits | pendent, not appressed to rachis, not or, rarely, weakly secund, straight, edges parallel, 1.5–3.5 cm × 1.2–2 mm; valves glabrous; ovules 28–44 per ovary; style 0.1–0.4 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight or curved, edges parallel, 3.5–6.5(–7.5) cm × 1–1.6 mm; valves glabrous; ovules 30–72 per ovary; style 0.1–1 mm. |
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Seeds | uniseriate, 0.9–1.2 × 0.6–1 mm; wing often continuous, 0.07–0.1 mm wide. |
uniseriate, 1–1.2 × 0.8–1 mm; wing continuous, ca. 0.1 mm wide. |
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Boechera oxylobula |
Boechera fernaldiana |
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Phenology | Flowering May–Jul. | |||||
Habitat | Cliffs, rocky slopes, gravelly soil in sagebrush and open conifer forests | |||||
Elevation | 2100-3600 m (6900-11800 ft) | |||||
Distribution |
CO |
CO; NV; UT |
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Discussion | Most of the collections assigned here have been called Arabis (Boechera) demissa by other authors (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b). Because the holotype of A. demissa is identical to B. oxylobula in nearly every way (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison), we treat them as conspecific. Boechera oxylobula is restricted to Garfield, Gunnison, Hinsdale, Lake, Mineral, Park, and Saguache counties in central Colorado. The taxon traditionally treated as A. (Boechera) demissa var. languida is here recognized as an apomictic species of hybrid origin (see Windham and Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Taxa treated here as subspecies of Boechera fernaldiana share many morphological traits, and most recent authors (R. C. Rollins 1993; N. H. Holmgren 2005b) did not recognize them as distinct. They are consistently separated by the characters listed below, show some degree of molecular divergence (C. D. Bailey et al., unpubl.), and their ranges are separated by ca. 500 km. They clearly represent genetically isolated population systems that warrant taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 391. | FNA vol. 7, p. 378. | ||||
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera | ||||
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Subordinate taxa | ||||||
Synonyms | Arabis oxylobula, Arabis aprica, Arabis demissa, Arabis rugocarpa, B. demissa | Arabis fernaldiana, Arabis canescens var. stylosa, Arabis fernaldiana var. stylosa | ||||
Name authority | (Greene) W. A. Weber: Phytologia 51: 370. (1982) | (Rollins) W. A. Weber: Phytologia 51: 370. (1982) | ||||
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