Boechera howellii |
Boechera pendulocarpa |
|
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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress |
danglepod rockcress, drop-seed rock-cress, earless suncress, Holboell's rockcress |
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Habit | Perennials; long-lived; sexual; caudex usually woody. | Perennials; short- to long-lived; sexual; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–3 dm, densely pubescent proximally, trichomes simple and stalked, 2–4-rayed, 0.1–0.4 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous. |
blade narrowly oblanceolate, 1.5–4(–5) mm wide, margins entire, often ciliate along petiole, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.08–0.2 mm. |
Cauline leaves | 2–4, not concealing stem; blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally. |
6–17, sometimes concealing stem proximally; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 2–5-flowered, unbranched. |
4–11-flowered, unbranched. |
Flowers | erect to ascending at anthesis; sepals glabrous; petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals white or lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | ascending, straight, 4–10 mm, glabrous. |
arched, gently recurved, 3–8(–12) mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm; valves glabrous; ovules 10–20 per ovary; style 0.05–0.3 mm. |
pendent, not appressed to rachis, not secund, usually straight, edges parallel, (2–)2.5–3.8 cm × 1.5–2.2 mm; valves glabrous; ovules 66–92 per ovary; style 0.3–0.5 mm. |
Seeds | uniseriate, 3–6 × 2–4 mm; wing continuous, 1.3–2.5 mm wide. |
sub-biseriate, 1–1.2 × 0.7–0.9 mm; wing distal, 0.05–0.1 mm wide. |
2n | = 14. |
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Boechera howellii |
Boechera pendulocarpa |
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Phenology | Flowering Jun–Aug. | Flowering Apr–Jul. |
Habitat | Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats | Rock outcrops and gravelly slopes in sagebrush, open conifer forests, subalpine meadows |
Elevation | 1800-3800 m (5900-12500 ft) | 1000-3300 m (3300-10800 ft) |
Distribution |
CA; NV; OR
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CA; ID; MT; NV; OR; UT; WY; BC; YT
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Discussion | Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993), B. pendulocarpa is easily distinguished from that species by having simple and 2–4-rayed (versus 4–8-rayed) trichomes proximally on stems, cauline leaves without auricles, fruiting pedicels gently (versus sharply) recurved, and shorter (2–)2.5–3.8 (versus 3.5–6.5) cm, non-secund fruits. The two taxa have allopatric distributions, with B. pendulocarpa found in the mountains of western North America and B. holboellii apparently confined to Greenland. Recent use of the name A. (Boechera) exilis for this taxon (e.g., G. A. Mulligan 1996; R. D. Dorn 2001; N. H. Holmgren 2005b) is based on misinterpretation of the type (M. D. Windham and I. A. Al-Shehbaz 2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 384. | FNA vol. 7, p. 395. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata | Arabis pendulocarpa, Arabis holboellii var. pendulocarpa, B. holboellii var. pendulocarpa |
Name authority | (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) | (A. Nelson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 77. (2006) |
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