Boechera howellii |
Boechera pallidifolia |
|
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Howell's flat-seed rock-cress, Howell's rockcress, pioneer rock-cress |
Gunnison County rockcress |
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Habit | Perennials; long-lived; sexual; caudex usually woody. | Perennials; short-lived; sexual; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–2(–3) dm, glabrous throughout. |
usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, (0.5–)1.5–4 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–7 mm wide, margins entire, ciliate proximally, trichomes (simple mixed with a few short-stalked, 2-rayed ones) 0.1–0.4 mm, surfaces glabrous. |
blade oblanceolate to obovate, 5–11(–13) mm wide, margins shallowly dentate or, sometimes, entire, often ciliate along petiole, trichomes (simple and 2-rayed), surfaces moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
Cauline leaves | 2–4, not concealing stem; blade auricles (0–)0.2–1 mm, surfaces of distalmost leaves usually glabrous, rarely ciliate proximally. |
3–8(–11), not concealing stem; blade auricles (0.5–)1–2 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 2–5-flowered, unbranched. |
4–15(–20)-flowered, usually unbranched. |
Flowers | erect to ascending at anthesis; sepals glabrous; petals white to dark lavender, 4–8 × 1–2.5 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals usually lavender, rarely whitish, 5–9 × 1.2–2.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | ascending, straight, 4–10 mm, glabrous. |
ascending to divaricate-ascending, straight or, sometimes, recurved, 7–15 mm, glabrous or sparsely pubescent. |
Fruits | suberect to ascending, usually not appressed to rachis, not secund, straight to curved, edges often somewhat undulate (not parallel), 2.5–6.5 cm × 3–7 mm; valves glabrous; ovules 10–20 per ovary; style 0.05–0.3 mm. |
usually ascending, rarely horizontal, not appressed to rachis, not secund, straight to curved, edges parallel, 2.5–5(–6.5) cm × 1–2 mm; valves glabrous; ovules 44–62 per ovary; style 0.05–0.4 mm. |
Seeds | uniseriate, 3–6 × 2–4 mm; wing continuous, 1.3–2.5 mm wide. |
uniseriate, 1.1–1.3 × 1–1.1 mm; wing nearly continuous, 0.07–0.15 mm wide. |
Boechera howellii |
Boechera pallidifolia |
|
Phenology | Flowering Jun–Aug. | Flowering Apr–Jun. |
Habitat | Rock outcrops, talus slopes and gravelly soil in alpine and subalpine habitats | Rocky slopes and sandy soil in pinyon-juniper, sagebrush communities |
Elevation | 1800-3800 m (5900-12500 ft) | 1600-2500 m (5200-8200 ft) |
Distribution |
CA; NV; OR
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AZ; CO; NM; UT; WY
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Discussion | Boechera howellii usually is subsumed under B. platysperma, but the two taxa are distinct (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although Boechera pallidifolia was originally described as endemic to west-central Colorado, recent studies favor a broader circumscription that includes sexual populations previously assigned to Arabis (Boechera) selbyi (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). The species has hybridized with nearly every sexual diploid within its range (including B. crandallii, B. fendleri, B. formosa, and B. pendulina), producing a confusing array of apomictic triploids including the type specimen of A. selbyi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 384. | FNA vol. 7, p. 392. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis howellii, Arabis conferta, Arabis inamoena var. acutata, Arabis platyloba, Arabis platysperma var. howellii, Arabis platysperma var. imparata | Arabis pallidifolia, Arabis thompsonii, B. thompsonii |
Name authority | (S. Watson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 69. (2006) | (Rollins) W. A. Weber: Phytologia 79: 65. (1996) |
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