Boechera holboellii |
Boechera williamsii |
|
---|---|---|
|
|
|
Habit | Biennials or perennials; short-lived; apomictic or sexual; caudex present or absent. | Perennials; short-lived; sexual; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6.7 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
Basal leaves | blade oblanceolate, 2–5 mm wide, margins entire, sometimes ciliate near petiole base, trichomes (usually simple), to 0.7 mm, surfaces sparsely to moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
Cauline leaves | 7–40, often concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 10–60-flowered, usually unbranched. |
5–17-flowered, unbranched. |
Flowers | divaricate to pendent at anthesis; sepals pubescent; petals white, 4–8 × 1–2 mm, glabrous; pollen ellipsoid or spheroid. |
erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | descending to reflexed, most abruptly recurved near base, 5–10 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
erect, straight, 2–4(–7) mm, glabrous. |
Fruits | descending to reflexed, not appressed to rachis, strongly secund, often curved, edges parallel, 0.35–0.65 cm × (2–)2.2–2.5 mm; valves glabrous; ovules 80–102 per ovary. |
erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate to sub-biseriate, 1.4–1.7 × 1–1.2 mm; wing continuous, to 0.2 mm wide. |
sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
2n | = 14, 21. |
|
Boechera holboellii |
Boechera williamsii |
|
Phenology | Flowering Jul–Aug. | Flowering May–Jun. |
Habitat | Rocky slopes and gravelly soil near coast | Gravelly soil in sagebrush-grassland communities |
Elevation | 100-300 m (300-1000 ft) | 2300-2800 m (7500-9200 ft) |
Distribution |
Greenland |
WY |
Discussion | Both sexual diploid and apomictic triploid collections of Boechera holboellii are known; available data suggest that the triploid is an autopolyploid derivative of the sexual diploid but further study is needed. In the narrow circumscription adopted here, it appears to be restricted to Greenland (see discussion of B. quebecensis in M. D. Windham and I. A. Al-Shehbaz 2007b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 384. | FNA vol. 7, p. 411. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis holboellii, Arabis holboellii var. tenuis, B. tenuis | Arabis williamsii |
Name authority | (Hornemann) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) |
Web links |