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Habit Biennials or perennials; short-lived; apomictic or sexual; caudex present or absent. Perennials; short-lived; sexual; caudex not woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6.7 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout.

Basal leaves

blade oblanceolate, 2–5 mm wide, margins entire, sometimes ciliate near petiole base, trichomes (usually simple), to 0.7 mm, surfaces sparsely to moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm.

blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm.

Cauline leaves

7–40, often concealing stem proximally;

blade auricles 1–4 mm, surfaces of distalmost leaves glabrous or sparsely pubescent.

6–11, sometimes concealing stem proximally;

blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous.

Racemes

10–60-flowered, usually unbranched.

5–17-flowered, unbranched.

Flowers

divaricate to pendent at anthesis;

sepals pubescent;

petals white, 4–8 × 1–2 mm, glabrous;

pollen ellipsoid or spheroid.

erect at anthesis;

sepals glabrous;

petals white, 3–4 × 0.5–0.8 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

descending to reflexed, most abruptly recurved near base, 5–10 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

erect, straight, 2–4(–7) mm, glabrous.

Fruits

descending to reflexed, not appressed to rachis, strongly secund, often curved, edges parallel, 0.35–0.65 cm × (2–)2.2–2.5 mm;

valves glabrous;

ovules 80–102 per ovary.

erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm;

valves glabrous;

ovules 60–96 per ovary;

style 0.05–0.2 mm.

Seeds

uniseriate to sub-biseriate, 1.4–1.7 × 1–1.2 mm;

wing continuous, to 0.2 mm wide.

sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally.

2n

= 14, 21.

Boechera holboellii

Boechera williamsii

Phenology Flowering Jul–Aug. Flowering May–Jun.
Habitat Rocky slopes and gravelly soil near coast Gravelly soil in sagebrush-grassland communities
Elevation 100-300 m (300-1000 ft) 2300-2800 m (7500-9200 ft)
Distribution
from USDA
Greenland
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Both sexual diploid and apomictic triploid collections of Boechera holboellii are known; available data suggest that the triploid is an autopolyploid derivative of the sexual diploid but further study is needed. In the narrow circumscription adopted here, it appears to be restricted to Greenland (see discussion of B. quebecensis in M. D. Windham and I. A. Al-Shehbaz 2007b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 384. FNA vol. 7, p. 411.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. xylopoda, B. yorkii
Synonyms Arabis holboellii, Arabis holboellii var. tenuis, B. tenuis Arabis williamsii
Name authority (Hornemann) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001)
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