Boechera holboellii |
Boechera rigidissima |
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Trinity Mountain rockcress |
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Habit | Biennials or perennials; short-lived; apomictic or sexual; caudex present or absent. | Perennials; long-lived; apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6.7 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, slightly elevated on woody base or from near ground surface, 2–6 dm, pubescent proximally, trichomes usually subsessile, 2- or 3-rayed, rarely simple, 0.25–0.4 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 2–5 mm wide, margins entire, sometimes ciliate near petiole base, trichomes (usually simple), to 0.7 mm, surfaces sparsely to moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
blade oblanceolate, 2–6 mm wide, margins entire, ciliate proximally, trichomes to 0.7 mm, surfaces sparsely pubescent, trichomes subsessile to short-stalked, 2–5-rayed, 0.1–0.4 mm. |
Cauline leaves | 7–40, often concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
5–10, rarely concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves glabrous. |
Racemes | 10–60-flowered, usually unbranched. |
5–16-flowered, usually unbranched. |
Flowers | divaricate to pendent at anthesis; sepals pubescent; petals white, 4–8 × 1–2 mm, glabrous; pollen ellipsoid or spheroid. |
ascending at anthesis; sepals glabrous; petals purple, 6–8 × 1.5–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | descending to reflexed, most abruptly recurved near base, 5–10 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
divaricate-ascending, straight, 4–10 mm, glabrous. |
Fruits | descending to reflexed, not appressed to rachis, strongly secund, often curved, edges parallel, 0.35–0.65 cm × (2–)2.2–2.5 mm; valves glabrous; ovules 80–102 per ovary. |
divaricate-ascending, not appressed to rachis, not secund, straight, edges somewhat undulate (not parallel), 4–6.5(–7.6) cm × 2.5–3.5 mm; valves glabrous; ovules 24–54 per ovary; style 0.2–0.8 mm. |
Seeds | uniseriate to sub-biseriate, 1.4–1.7 × 1–1.2 mm; wing continuous, to 0.2 mm wide. |
uniseriate, 2.5–3.2 × 1.8–2.5 mm; wing distal, 0.3–1 mm wide. |
2n | = 14, 21. |
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Boechera holboellii |
Boechera rigidissima |
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Phenology | Flowering Jul–Aug. | Flowering Jul–Aug. |
Habitat | Rocky slopes and gravelly soil near coast | Rocky places in open conifer forests |
Elevation | 100-300 m (300-1000 ft) | 1800-2000 m (5900-6600 ft) |
Distribution |
Greenland |
CA; NV |
Discussion | Both sexual diploid and apomictic triploid collections of Boechera holboellii are known; available data suggest that the triploid is an autopolyploid derivative of the sexual diploid but further study is needed. In the narrow circumscription adopted here, it appears to be restricted to Greenland (see discussion of B. quebecensis in M. D. Windham and I. A. Al-Shehbaz 2007b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Plants assigned to Boechera rigidissima appear to represent a series of apomictic hybrids, possibly involving B. stricta and B. suffrutescens. The two varieties recognized by Rollins are well isolated geographically (in Trinity County, California, and Washoe County, Nevada, respectively) and show enough morphological divergence to suggest that they represent distinct origins, if not different combinations of parental genomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 384. | FNA vol. 7, p. 403. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis holboellii, Arabis holboellii var. tenuis, B. tenuis | Arabis rigidissima, Arabis rigidissima var. demota |
Name authority | (Hornemann) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) | (Rollins) Al-Shehbaz: Novon 13: 389. (2003) |
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