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Pinzl's Rock cress

Habit Biennials or perennials; short-lived; apomictic or sexual; caudex present or absent. Perennials; long-lived; apomictic; caudex usually woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6.7 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally.

1–3 per caudex branch, arising from margin of rosette near ground surface, 0.4–1.6 dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.05–0.2 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade oblanceolate, 2–5 mm wide, margins entire, sometimes ciliate near petiole base, trichomes (usually simple), to 0.7 mm, surfaces sparsely to moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm.

(petiole glabrous);

blade linear-oblanceolate, 1–3 mm wide, margins entire, rarely ciliate proximally, trichomes to 0.5 mm, surfaces densely pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.2 mm.

Cauline leaves

7–40, often concealing stem proximally;

blade auricles 1–4 mm, surfaces of distalmost leaves glabrous or sparsely pubescent.

3–6, not concealing stem;

blade auricles 0(–0.3) mm, surfaces of distalmost leaves sparsely to moderately pubescent.

Racemes

10–60-flowered, usually unbranched.

5–8-flowered, unbranched.

Flowers

divaricate to pendent at anthesis;

sepals pubescent;

petals white, 4–8 × 1–2 mm, glabrous;

pollen ellipsoid or spheroid.

ascending at anthesis;

sepals pubescent;

petals purple, 4–5 × 1–1.5 mm, glabrous;

pollen spheroid.

Fruiting pedicels

descending to reflexed, most abruptly recurved near base, 5–10 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

ascending to divaricate-ascending, straight, 2–6 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

Fruits

descending to reflexed, not appressed to rachis, strongly secund, often curved, edges parallel, 0.35–0.65 cm × (2–)2.2–2.5 mm;

valves glabrous;

ovules 80–102 per ovary.

ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 2.5–4.8 cm × 2.5–3.2 mm;

valves glabrous;

ovules 26–34 per ovary;

style 0.1–0.3 mm.

Seeds

uniseriate to sub-biseriate, 1.4–1.7 × 1–1.2 mm;

wing continuous, to 0.2 mm wide.

uniseriate, 2.5–3.5 × 1.5–2.5 mm;

wing continuous, 0.2–0.9 mm wide.

2n

= 14, 21.

Boechera holboellii

Boechera pinzliae

Phenology Flowering Jul–Aug. Flowering Jul.
Habitat Rocky slopes and gravelly soil near coast Gravelly granitic soils in alpine and subalpine areas
Elevation 100-300 m (300-1000 ft) 3000-3400 m (9800-11200 ft)
Distribution
from USDA
Greenland
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Both sexual diploid and apomictic triploid collections of Boechera holboellii are known; available data suggest that the triploid is an autopolyploid derivative of the sexual diploid but further study is needed. In the narrow circumscription adopted here, it appears to be restricted to Greenland (see discussion of B. quebecensis in M. D. Windham and I. A. Al-Shehbaz 2007b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Morphological evidence suggests that Boechera pinzliae is an apomictic hybrid that contains at least one genome derived from B. platysperma; the other parent has not yet been determined. It is most similar to B. elkoensis, from which it differs in having sparsely pubescent (versus glabrous) sepals and basal leaves with stalked (versus sessile or subsessile) trichomes. Boechera pinzliae is known only from the White Mountains in extreme western Nevada (Esmeralda County) and east-central California (Mono County).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 384. FNA vol. 7, p. 397.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis holboellii, Arabis holboellii var. tenuis, B. tenuis Arabis pinzliae
Name authority (Hornemann) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) (Rollins) Al-Shehbaz: Novon 13: 388. (2003)
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