Boechera holboellii |
Boechera pinzliae |
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Pinzl's Rock cress |
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Habit | Biennials or perennials; short-lived; apomictic or sexual; caudex present or absent. | Perennials; long-lived; apomictic; caudex usually woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6.7 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
1–3 per caudex branch, arising from margin of rosette near ground surface, 0.4–1.6 dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.05–0.2 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 2–5 mm wide, margins entire, sometimes ciliate near petiole base, trichomes (usually simple), to 0.7 mm, surfaces sparsely to moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
(petiole glabrous); blade linear-oblanceolate, 1–3 mm wide, margins entire, rarely ciliate proximally, trichomes to 0.5 mm, surfaces densely pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.2 mm. |
Cauline leaves | 7–40, often concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
3–6, not concealing stem; blade auricles 0(–0.3) mm, surfaces of distalmost leaves sparsely to moderately pubescent. |
Racemes | 10–60-flowered, usually unbranched. |
5–8-flowered, unbranched. |
Flowers | divaricate to pendent at anthesis; sepals pubescent; petals white, 4–8 × 1–2 mm, glabrous; pollen ellipsoid or spheroid. |
ascending at anthesis; sepals pubescent; petals purple, 4–5 × 1–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | descending to reflexed, most abruptly recurved near base, 5–10 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
ascending to divaricate-ascending, straight, 2–6 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | descending to reflexed, not appressed to rachis, strongly secund, often curved, edges parallel, 0.35–0.65 cm × (2–)2.2–2.5 mm; valves glabrous; ovules 80–102 per ovary. |
ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 2.5–4.8 cm × 2.5–3.2 mm; valves glabrous; ovules 26–34 per ovary; style 0.1–0.3 mm. |
Seeds | uniseriate to sub-biseriate, 1.4–1.7 × 1–1.2 mm; wing continuous, to 0.2 mm wide. |
uniseriate, 2.5–3.5 × 1.5–2.5 mm; wing continuous, 0.2–0.9 mm wide. |
2n | = 14, 21. |
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Boechera holboellii |
Boechera pinzliae |
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Phenology | Flowering Jul–Aug. | Flowering Jul. |
Habitat | Rocky slopes and gravelly soil near coast | Gravelly granitic soils in alpine and subalpine areas |
Elevation | 100-300 m (300-1000 ft) | 3000-3400 m (9800-11200 ft) |
Distribution |
Greenland |
CA; NV |
Discussion | Both sexual diploid and apomictic triploid collections of Boechera holboellii are known; available data suggest that the triploid is an autopolyploid derivative of the sexual diploid but further study is needed. In the narrow circumscription adopted here, it appears to be restricted to Greenland (see discussion of B. quebecensis in M. D. Windham and I. A. Al-Shehbaz 2007b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Morphological evidence suggests that Boechera pinzliae is an apomictic hybrid that contains at least one genome derived from B. platysperma; the other parent has not yet been determined. It is most similar to B. elkoensis, from which it differs in having sparsely pubescent (versus glabrous) sepals and basal leaves with stalked (versus sessile or subsessile) trichomes. Boechera pinzliae is known only from the White Mountains in extreme western Nevada (Esmeralda County) and east-central California (Mono County). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 384. | FNA vol. 7, p. 397. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis holboellii, Arabis holboellii var. tenuis, B. tenuis | Arabis pinzliae |
Name authority | (Hornemann) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) | (Rollins) Al-Shehbaz: Novon 13: 388. (2003) |
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