Boechera harrisonii |
Boechera pendulocarpa |
|
---|---|---|
|
danglepod rockcress, drop-seed rock-cress, earless suncress, Holboell's rockcress |
|
Habit | Perennials; long-lived; (some-what cespitose); apomictic; caudex often woody. | Perennials; short- to long-lived; sexual; caudex usually not woody. |
Stems | usually 3–7 per caudex branch, arising from margin of rosette near ground surface, 0.5–2.5 dm, usually sparsely pubescent proximally, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.06–0.2 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.6–3 dm, densely pubescent proximally, trichomes simple and stalked, 2–4-rayed, 0.1–0.4 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 2–4 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 0.5 mm, surfaces sparsely pubescent, trichomes short-stalked, 2–5-rayed, 0.1–0.25 mm. |
blade narrowly oblanceolate, 1.5–4(–5) mm wide, margins entire, often ciliate along petiole, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.08–0.2 mm. |
Cauline leaves | 3–7, not concealing stem; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
6–17, sometimes concealing stem proximally; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 5–12-flowered, usually unbranched. |
4–11-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 5–7.5 × 0.8–1.5 mm, glabrous; pollen spheroid. |
ascending at anthesis; sepals pubescent; petals white or lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | divaricate-ascending, straight, 8–12 mm, glabrous. |
arched, gently recurved, 3–8(–12) mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | usually divaricate-ascending, rarely nearly horizontal, not appressed to rachis, not secund, curved to straight, edges parallel, 3–4.7 cm × 1–1.5 mm; valves glabrous; ovules 64–86 per ovary; style 0.2–0.5 mm. |
pendent, not appressed to rachis, not secund, usually straight, edges parallel, (2–)2.5–3.8 cm × 1.5–2.2 mm; valves glabrous; ovules 66–92 per ovary; style 0.3–0.5 mm. |
Seeds | uniseriate, 1.1–1.3 × 0.8–1.1 mm; wing continuous, 0.08–0.1 mm wide. |
sub-biseriate, 1–1.2 × 0.7–0.9 mm; wing distal, 0.05–0.1 mm wide. |
2n | = 14. |
|
Boechera harrisonii |
Boechera pendulocarpa |
|
Phenology | Flowering May–Jun. | Flowering Apr–Jul. |
Habitat | Limestone and quartzite cliffs | Rock outcrops and gravelly slopes in sagebrush, open conifer forests, subalpine meadows |
Elevation | 1500-1600 m (4900-5200 ft) | 1000-3300 m (3300-10800 ft) |
Distribution |
UT |
CA; ID; MT; NV; OR; UT; WY; BC; YT
|
Discussion | Morphological evidence suggests that Boechera harrisonii is an apomictic species that arose through hybridization between B. microphylla and B. perennans (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). Boechera harrisonii is known only from Utah and Wasatch counties in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993), B. pendulocarpa is easily distinguished from that species by having simple and 2–4-rayed (versus 4–8-rayed) trichomes proximally on stems, cauline leaves without auricles, fruiting pedicels gently (versus sharply) recurved, and shorter (2–)2.5–3.8 (versus 3.5–6.5) cm, non-secund fruits. The two taxa have allopatric distributions, with B. pendulocarpa found in the mountains of western North America and B. holboellii apparently confined to Greenland. Recent use of the name A. (Boechera) exilis for this taxon (e.g., G. A. Mulligan 1996; R. D. Dorn 2001; N. H. Holmgren 2005b) is based on misinterpretation of the type (M. D. Windham and I. A. Al-Shehbaz 2006). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 383. | FNA vol. 7, p. 395. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis harrisonii, B. microphylla var. harrisonii | Arabis pendulocarpa, Arabis holboellii var. pendulocarpa, B. holboellii var. pendulocarpa |
Name authority | (S. L. Welsh) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) | (A. Nelson) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 77. (2006) |
Web links |
|