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Graham's rockcress, Graham's suncress

Habit Perennials; long-lived; (some-what cespitose); apomictic; caudex often woody. Biennials or perennials; short-lived; apomictic; caudex present or absent.
Stems

usually 3–7 per caudex branch, arising from margin of rosette near ground surface, 0.5–2.5 dm, usually sparsely pubescent proximally, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.06–0.2 mm, glabrous distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, (1.2–)2–12 dm, glabrous or pubescent proximally, trichomes simple and sessile or subsessile, 2 or 3(–6)-rayed, to 1 mm, glabrous distally.

Basal leaves

blade narrowly oblanceolate, 2–4 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 0.5 mm, surfaces sparsely pubescent, trichomes short-stalked, 2–5-rayed, 0.1–0.25 mm.

blade oblanceolate, 1.5–10(–20) mm wide, margins entire or dentate, ciliate proximally, trichomes (simple), to 1 mm, surfaces sparsely to densely pubescent, trichomes sessile or subsessile, 2–4(–7)-rayed, 0.2–0.6 mm.

Cauline leaves

3–7, not concealing stem;

blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous.

(10–)13–52, often concealing stem proximally;

blade auricles 1–5 mm, surfaces of distalmost leaves glabrous.

Racemes

5–12-flowered, usually unbranched.

16–88-flowered, usually unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals lavender, 5–7.5 × 0.8–1.5 mm, glabrous;

pollen spheroid.

divaricate at anthesis;

sepals glabrous or pubescent;

petals usually white, rarely lavender, 5.5–8 × 1–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

divaricate-ascending, straight, 8–12 mm, glabrous.

divaricate-ascending to descending, usually gently curved downward, rarely straight, 5–22(–30) mm, glabrous.

Fruits

usually divaricate-ascending, rarely nearly horizontal, not appressed to rachis, not secund, curved to straight, edges parallel, 3–4.7 cm × 1–1.5 mm;

valves glabrous;

ovules 64–86 per ovary;

style 0.2–0.5 mm.

divaricate-ascending to pendent, not appressed to rachis, not secund, straight or slightly curved, edges parallel, 3.5–9 cm × 1–1.8 mm;

valves glabrous;

ovules 84–146 per ovary;

style 0.05–0.5 mm.

Seeds

uniseriate, 1.1–1.3 × 0.8–1.1 mm;

wing continuous, 0.08–0.1 mm wide.

uniseriate to sub-biseriate, 1–1.8 × 0.8–1.2 mm;

wing continuous, 0.1–0.25 mm wide.

2n

= 21.

Boechera harrisonii

Boechera grahamii

Phenology Flowering May–Jun. Flowering May–Aug.
Habitat Limestone and quartzite cliffs Rocky slopes and sandy soil in prairies and open forests
Elevation 1500-1600 m (4900-5200 ft) 100-3200 m (300-10500 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
AK; CO; MI; MN; MT; ND; NY; SD; VT; WA; WI; WY; AB; BC; MB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Morphological evidence suggests that Boechera harrisonii is an apomictic species that arose through hybridization between B. microphylla and B. perennans (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). Boechera harrisonii is known only from Utah and Wasatch counties in north-central Utah.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera grahamii is an apomictic species that arose through hybridization between B. collinsii and B. stricta. Previous authors have assigned these specimens to Arabis (Boechera) divaricarpa (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison); if these taxa are treated as conspecific, the name B. grahamii has priority.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 383. FNA vol. 7, p. 382.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis harrisonii, B. microphylla var. harrisonii Turritis grahamii, Arabis boivinii, Arabis bourgovii, Arabis confinis, Arabis confinis var. brachycarpa, Arabis dacotica, Arabis divaricarpa var. dacotica, Arabis divaricarpa var. hemicylindrica, Arabis divaricarpa var. stenocarpa, Arabis drummondii var. brachycarpa, Arabis holboellii var. brachycarpa, B. brachycarpa, Turritis brachycarpa
Name authority (S. L. Welsh) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) (Lehmann) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 241. (2007)
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