Boechera harrisonii |
Boechera fructicosa |
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Habit | Perennials; long-lived; (some-what cespitose); apomictic; caudex often woody. | Perennials; short-lived; apomictic; caudex usually not woody. |
Stems | usually 3–7 per caudex branch, arising from margin of rosette near ground surface, 0.5–2.5 dm, usually sparsely pubescent proximally, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.06–0.2 mm, glabrous distally. |
3 to many per caudex branch, arising from margin of rosette near ground surface, 3.5–6 dm, pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.2–0.3 mm, glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 2–4 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 0.5 mm, surfaces sparsely pubescent, trichomes short-stalked, 2–5-rayed, 0.1–0.25 mm. |
blade oblanceolate, 3–7 mm wide, margins dentate, ciliate proximally, trichomes (simple), to 0.3 mm, surfaces pubescent, trichomes short-stalked, 4–8-rayed, 0.05–0.2 mm. |
Cauline leaves | 3–7, not concealing stem; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
5–10, not concealing stem; blade auricles 0.5–1 mm, surfaces of distalmost leaves glabrous. |
Racemes | 5–12-flowered, usually unbranched. |
10–18-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 5–7.5 × 0.8–1.5 mm, glabrous; pollen spheroid. |
divaricate-ascending at anthesis; sepals pubescent; petals lavender, 4–6 × 1–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | divaricate-ascending, straight, 8–12 mm, glabrous. |
divaricate-descending, straight, 5–10 mm, glabrous. |
Fruits | usually divaricate-ascending, rarely nearly horizontal, not appressed to rachis, not secund, curved to straight, edges parallel, 3–4.7 cm × 1–1.5 mm; valves glabrous; ovules 64–86 per ovary; style 0.2–0.5 mm. |
divaricate-descending, not appressed to rachis, not secund, curved, edges parallel, 4–6 cm × 1.5–1.8 mm; valves glabrous; ovules 60–70 per ovary; style 0.1–0.4 mm. |
Seeds | uniseriate, 1.1–1.3 × 0.8–1.1 mm; wing continuous, 0.08–0.1 mm wide. |
uniseriate, 1.5–2 × 1–1.3 mm; wing distal, ca. 0.1 mm wide. |
Boechera harrisonii |
Boechera fructicosa |
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Phenology | Flowering May–Jun. | Flowering Jun–Jul. |
Habitat | Limestone and quartzite cliffs | Dry, disturbed soil |
Elevation | 1500-1600 m (4900-5200 ft) | ca. 2000 m (ca. 6600 ft) |
Distribution |
UT |
WY |
Discussion | Morphological evidence suggests that Boechera harrisonii is an apomictic species that arose through hybridization between B. microphylla and B. perennans (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). Boechera harrisonii is known only from Utah and Wasatch counties in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera fructicosa is known only from the type collection (A. & E. Nelson 5681) taken near Undine Falls in Park County. Multiple plants are represented (comprising the holotype and four isotypes), though one specimen with the same number at RM is B. stricta. Although the exact parentage of this apomictic hybrid remains in doubt, it is virtually certain that B. microphylla contributed at least one genome. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 383. | FNA vol. 7, p. 379. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis harrisonii, B. microphylla var. harrisonii | Arabis fructicosa |
Name authority | (S. L. Welsh) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) | (A. Nelson) Al-Shehbaz: Novon 13: 385. (2003) |
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