Boechera harrisonii |
Boechera fernaldiana |
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Habit | Perennials; long-lived; (some-what cespitose); apomictic; caudex often woody. | Perennials; long-lived; (cespitose); sexual; caudex somewhat woody (sometimes with persistent, crowded leaf bases). | ||||
Stems | usually 3–7 per caudex branch, arising from margin of rosette near ground surface, 0.5–2.5 dm, usually sparsely pubescent proximally, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.06–0.2 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette, near ground surface or slightly elevated on woody base, 1–3.8 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.04–0.1 mm, sometimes mixed with stalked, 2-rayed ones, to 0.5 mm, usually sparsely pubescent distally. |
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Basal leaves | blade narrowly oblanceolate, 2–4 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 0.5 mm, surfaces sparsely pubescent, trichomes short-stalked, 2–5-rayed, 0.1–0.25 mm. |
blade narrowly oblanceolate, 1–4 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.04–0.1 mm. |
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Cauline leaves | 3–7, not concealing stem; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
5–10, not concealing stem; blade auricles 0.3–2 mm, surfaces of distalmost leaves usually sparsely pubescent. |
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Racemes | 5–12-flowered, usually unbranched. |
10–20-flowered, usually unbranched. |
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Flowers | ascending at anthesis; sepals pubescent; petals lavender, 5–7.5 × 0.8–1.5 mm, glabrous; pollen spheroid. |
ascending at anthesis; sepals (purplish or greenish) glabrous or sparsely or moderately pubescent; petals white or purple to lavender, 7–12 × 1.5–4 mm, glabrous; pollen ellipsoid. |
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Fruiting pedicels | divaricate-ascending, straight, 8–12 mm, glabrous. |
divaricate-ascending, straight, 5–20 mm, sparsely pubescent, trichomes appressed, branched. |
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Fruits | usually divaricate-ascending, rarely nearly horizontal, not appressed to rachis, not secund, curved to straight, edges parallel, 3–4.7 cm × 1–1.5 mm; valves glabrous; ovules 64–86 per ovary; style 0.2–0.5 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight or curved, edges parallel, 3.5–6.5(–7.5) cm × 1–1.6 mm; valves glabrous; ovules 30–72 per ovary; style 0.1–1 mm. |
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Seeds | uniseriate, 1.1–1.3 × 0.8–1.1 mm; wing continuous, 0.08–0.1 mm wide. |
uniseriate, 1–1.2 × 0.8–1 mm; wing continuous, ca. 0.1 mm wide. |
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Boechera harrisonii |
Boechera fernaldiana |
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Phenology | Flowering May–Jun. | |||||
Habitat | Limestone and quartzite cliffs | |||||
Elevation | 1500-1600 m (4900-5200 ft) | |||||
Distribution |
UT |
CO; NV; UT |
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Discussion | Morphological evidence suggests that Boechera harrisonii is an apomictic species that arose through hybridization between B. microphylla and B. perennans (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). Boechera harrisonii is known only from Utah and Wasatch counties in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Taxa treated here as subspecies of Boechera fernaldiana share many morphological traits, and most recent authors (R. C. Rollins 1993; N. H. Holmgren 2005b) did not recognize them as distinct. They are consistently separated by the characters listed below, show some degree of molecular divergence (C. D. Bailey et al., unpubl.), and their ranges are separated by ca. 500 km. They clearly represent genetically isolated population systems that warrant taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 383. | FNA vol. 7, p. 378. | ||||
Parent taxa | ||||||
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Synonyms | Arabis harrisonii, B. microphylla var. harrisonii | Arabis fernaldiana, Arabis canescens var. stylosa, Arabis fernaldiana var. stylosa | ||||
Name authority | (S. L. Welsh) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) | (Rollins) W. A. Weber: Phytologia 51: 370. (1982) | ||||
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