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Four Corners rockcress

Habit Perennials; long-lived; (some-what cespitose); apomictic; caudex often woody. Biennials or perennials; short-lived; apomictic; caudex present or absent.
Stems

usually 3–7 per caudex branch, arising from margin of rosette near ground surface, 0.5–2.5 dm, usually sparsely pubescent proximally, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.06–0.2 mm, glabrous distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–5 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.2–0.5 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade narrowly oblanceolate, 2–4 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 0.5 mm, surfaces sparsely pubescent, trichomes short-stalked, 2–5-rayed, 0.1–0.25 mm.

blade oblanceolate, 2–10 mm wide, margins usually dentate, sometimes ciliate near petiole base, trichomes (usually spurred), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.4 mm.

Cauline leaves

3–7, not concealing stem;

blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous.

15–36, sometimes concealing stem proximally;

blade auricles 1–3 mm, surfaces of distalmost leaves sparsely pubescent.

Racemes

5–12-flowered, usually unbranched.

20–55-flowered, usually unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals lavender, 5–7.5 × 0.8–1.5 mm, glabrous;

pollen spheroid.

divaricate-descending at anthesis;

sepals pubescent;

petals pale lavender, 5–8.5 × 1–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

divaricate-ascending, straight, 8–12 mm, glabrous.

reflexed to divaricate-descending, usually curved downward, 8–14 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched.

Fruits

usually divaricate-ascending, rarely nearly horizontal, not appressed to rachis, not secund, curved to straight, edges parallel, 3–4.7 cm × 1–1.5 mm;

valves glabrous;

ovules 64–86 per ovary;

style 0.2–0.5 mm.

reflexed to pendent, rarely appressed to rachis, not secund, straight or slightly curved, edges parallel, 4–6 cm × 1–2 mm;

valves glabrous;

ovules 100–128 per ovary;

style 0.05–0.5 mm.

Seeds

uniseriate, 1.1–1.3 × 0.8–1.1 mm;

wing continuous, 0.08–0.1 mm wide.

biseriate to sub-biseriate, mature seeds not seen.

Boechera harrisonii

Boechera consanguinea

Phenology Flowering May–Jun. Flowering May.
Habitat Limestone and quartzite cliffs Rocky slopes and sandy soil in ponderosa pine, pinyon-juniper, and sagebrush communities
Elevation 1500-1600 m (4900-5200 ft) 1900-2500 m (6200-8200 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Morphological evidence suggests that Boechera harrisonii is an apomictic species that arose through hybridization between B. microphylla and B. perennans (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). Boechera harrisonii is known only from Utah and Wasatch counties in north-central Utah.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera consanguinea is an apomictic species that arose through hybridization between B. fendleri and B. retrofracta. It is most similar to B. goodrichii, another apomictic hybrid involving B. retrofracta (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 383. FNA vol. 7, p. 371.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis harrisonii, B. microphylla var. harrisonii Arabis consanguinea, Arabis holboellii var. consanguinea
Name authority (S. L. Welsh) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 261. (2007)
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