Boechera fructicosa |
Boechera texana |
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Habit | Perennials; short-lived; apomictic; caudex usually not woody. | Perennials; long-lived; sexual; caudex woody. |
Stems | 3 to many per caudex branch, arising from margin of rosette near ground surface, 3.5–6 dm, pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.2–0.3 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette, often slightly elevated above ground surface on woody base, 2–5 dm, glabrous throughout. |
Basal leaves | blade oblanceolate, 3–7 mm wide, margins dentate, ciliate proximally, trichomes (simple), to 0.3 mm, surfaces pubescent, trichomes short-stalked, 4–8-rayed, 0.05–0.2 mm. |
blade oblanceolate, 5–12 mm wide, margins dentate, ciliate (at least proximally), trichomes (usually simple), 1–2 mm, surfaces moderately pubescent, trichomes short-stalked, 2–4-rayed, 0.4–0.6 mm. |
Cauline leaves | 5–10, not concealing stem; blade auricles 0.5–1 mm, surfaces of distalmost leaves glabrous. |
5–12, often concealing stem proximally; blade auricles 1–2 mm, surfaces distally glabrous. |
Racemes | 10–18-flowered, usually unbranched. |
10–35-flowered, usually unbranched. |
Flowers | divaricate-ascending at anthesis; sepals pubescent; petals lavender, 4–6 × 1–1.5 mm, glabrous; pollen spheroid. |
divaricate-ascending at anthesis; sepals glabrous; petals white to lavender, 5–8 × 1.5–2 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | divaricate-descending, straight, 5–10 mm, glabrous. |
horizontal to divaricate-ascending, curved or angled downward, 10–20 mm, glabrous. |
Fruits | divaricate-descending, not appressed to rachis, not secund, curved, edges parallel, 4–6 cm × 1.5–1.8 mm; valves glabrous; ovules 60–70 per ovary; style 0.1–0.4 mm. |
widely pendent, not appressed to rachis, not secund, curved to nearly straight, edges parallel, 3.5–5 cm × 2.5–3 mm; valves glabrous; ovules 80–130 per ovary; style 0.2–0.7 mm. |
Seeds | uniseriate, 1.5–2 × 1–1.3 mm; wing distal, ca. 0.1 mm wide. |
biseriate, 1.1–1.3 × 0.9–1 mm; wing distal or continuous, 0.1–0.15 mm wide. |
Boechera fructicosa |
Boechera texana |
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Phenology | Flowering Jun–Jul. | Flowering Apr. |
Habitat | Dry, disturbed soil | Rock outcrops and gravelly soils in desert grassland and evergreen woodlands |
Elevation | ca. 2000 m (ca. 6600 ft) | 1200-1700 m (3900-5600 ft) |
Distribution |
WY |
NM; TX |
Discussion | Boechera fructicosa is known only from the type collection (A. & E. Nelson 5681) taken near Undine Falls in Park County. Multiple plants are represented (comprising the holotype and four isotypes), though one specimen with the same number at RM is B. stricta. Although the exact parentage of this apomictic hybrid remains in doubt, it is virtually certain that B. microphylla contributed at least one genome. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera texana is a sexual diploid included within Arabis fendleri var. fendleri by R. C. Rollins (1993), but it appears to be sufficiently distinct to warrant recognition at species level (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). There is little or no geographic overlap between the two, with B. texana confined to western Texas and southern New Mexico and B. fendleri ranging from western New Mexico and the Four Corners region through northern Arizona to southern Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 379. | FNA vol. 7, p. 409. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis fructicosa | |
Name authority | (A. Nelson) Al-Shehbaz: Novon 13: 385. (2003) | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 85. (2006) |
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