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Tulare County rock cress

Habit Perennials; short-lived; apomictic; caudex usually not woody. Perennials; usually long-lived; sexual; caudex woody, (often with persistent, crowded leaf bases).
Stems

3 to many per caudex branch, arising from margin of rosette near ground surface, 3.5–6 dm, pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.2–0.3 mm, glabrous distally.

usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.2–0.8 dm, pubescent proximally, trichomes simple and short-stalked, 2- or 3-rayed, to 0.4 mm, glabrescent distally.

Basal leaves

blade oblanceolate, 3–7 mm wide, margins dentate, ciliate proximally, trichomes (simple), to 0.3 mm, surfaces pubescent, trichomes short-stalked, 4–8-rayed, 0.05–0.2 mm.

blade linear, 0.8–1.5 mm wide, margins entire, ciliate, trichomes (simple and spurred), to 0.8 mm, surfaces moderately pubescent, trichomes short-stalked, 2–4-rayed, 0.05–0.4 mm.

Cauline leaves

5–10, not concealing stem;

blade auricles 0.5–1 mm, surfaces of distalmost leaves glabrous.

2–4, not concealing stem;

blade auricles absent, surfaces of distalmost leaves pubescent.

Racemes

10–18-flowered, usually unbranched.

2–5-flowered, unbranched.

Flowers

divaricate-ascending at anthesis;

sepals pubescent;

petals lavender, 4–6 × 1–1.5 mm, glabrous;

pollen spheroid.

erect at anthesis;

sepals pubescent;

petals white, 3.5–5 × 0.7–1 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

divaricate-descending, straight, 5–10 mm, glabrous.

erect to ascending, straight, 2–7 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched.

Fruits

divaricate-descending, not appressed to rachis, not secund, curved, edges parallel, 4–6 cm × 1.5–1.8 mm;

valves glabrous;

ovules 60–70 per ovary;

style 0.1–0.4 mm.

erect to ascending, often appressed to rachis, not secund, straight, edges parallel, (1.3–)2–3.3 cm × 4–5 mm;

valves glabrous;

ovules 8–12 per ovary;

style 0.05–0.4 mm.

Seeds

uniseriate, 1.5–2 × 1–1.3 mm;

wing distal, ca. 0.1 mm wide.

uniseriate, 3–5 × 2.5–4.5 mm;

wing continuous, 0.8–2 mm wide distally.

Boechera fructicosa

Boechera pygmaea

Phenology Flowering Jun–Jul. Flowering May–Jul.
Habitat Dry, disturbed soil Barren flats of arkosic gravel
Elevation ca. 2000 m (ca. 6600 ft) 2400-3200 m (7900-10500 ft)
Distribution
from FNA
WY
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Boechera fructicosa is known only from the type collection (A. & E. Nelson 5681) taken near Undine Falls in Park County. Multiple plants are represented (comprising the holotype and four isotypes), though one specimen with the same number at RM is B. stricta. Although the exact parentage of this apomictic hybrid remains in doubt, it is virtually certain that B. microphylla contributed at least one genome.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Despite the distinctive morphology of Boechera pygmaea, there is evidence that it is capable of producing fertile hybrids with B. stricta. It is known only from Inyo and Tulare counties in the southern Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 379. FNA vol. 7, p. 401.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis fructicosa Arabis pygmaea
Name authority (A. Nelson) Al-Shehbaz: Novon 13: 385. (2003) (Rollins) Al-Shehbaz: Novon 13: 388. (2003)
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