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Coville's rock-cress

Habit Perennials; short-lived; apomictic; caudex usually not woody. Perennials; long-lived; apomictic; caudex woody.
Stems

3 to many per caudex branch, arising from margin of rosette near ground surface, 3.5–6 dm, pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.2–0.3 mm, glabrous distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.5–2.5 dm, glabrous throughout.

Basal leaves

blade oblanceolate, 3–7 mm wide, margins dentate, ciliate proximally, trichomes (simple), to 0.3 mm, surfaces pubescent, trichomes short-stalked, 4–8-rayed, 0.05–0.2 mm.

blade oblanceolate, 1.5–5 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.4 mm, and distally, trichomes short-stalked, 2 or 3 (or 4)-rayed, 0.15–0.4 mm, surfaces glabrous.

Cauline leaves

5–10, not concealing stem;

blade auricles 0.5–1 mm, surfaces of distalmost leaves glabrous.

2–7, not concealing stem;

blade with auricles 0.2–1 mm, auricles rarely absent, surfaces of distalmost leaves glabrous.

Racemes

10–18-flowered, usually unbranched.

2–9-flowered, unbranched.

Flowers

divaricate-ascending at anthesis;

sepals pubescent;

petals lavender, 4–6 × 1–1.5 mm, glabrous;

pollen spheroid.

ascending at anthesis;

sepals glabrous;

petals lavender to purplish, 5–6 × 1–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

divaricate-descending, straight, 5–10 mm, glabrous.

ascending to suberect, straight, 6–20 mm, glabrous.

Fruits

divaricate-descending, not appressed to rachis, not secund, curved, edges parallel, 4–6 cm × 1.5–1.8 mm;

valves glabrous;

ovules 60–70 per ovary;

style 0.1–0.4 mm.

ascending to suberect, rarely appressed to rachis, not secund, straight, edges slightly undulate (rarely parallel), 3.5–5 cm × 3–5 mm;

valves glabrous;

ovules 18–30 per ovary;

style 0.2–1 mm.

Seeds

uniseriate, 1.5–2 × 1–1.3 mm;

wing distal, ca. 0.1 mm wide.

uniseriate, 4–5 × 3–4 mm;

wing continuous, 1–1.5 mm wide.

Boechera fructicosa

Boechera covillei

Phenology Flowering Jun–Jul. Flowering Jul–Aug.
Habitat Dry, disturbed soil Rocky slopes in alpine meadows and open coniferous forests
Elevation ca. 2000 m (ca. 6600 ft) 2200-3500 m (7200-11500 ft)
Distribution
from FNA
WY
[BONAP county map]
from FNA
CA; NV; OR
[BONAP county map]
Discussion

Boechera fructicosa is known only from the type collection (A. & E. Nelson 5681) taken near Undine Falls in Park County. Multiple plants are represented (comprising the holotype and four isotypes), though one specimen with the same number at RM is B. stricta. Although the exact parentage of this apomictic hybrid remains in doubt, it is virtually certain that B. microphylla contributed at least one genome.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera covillei is an apomictic species that arose through hybridization between B. howellii and B. lyallii (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 379. FNA vol. 7, p. 371.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis fructicosa Arabis covillei, Arabis leibergii
Name authority (A. Nelson) Al-Shehbaz: Novon 13: 385. (2003) (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 237. (2007)
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