Boechera crandallii |
Boechera pinetorum |
|
---|---|---|
Crandall's rockcress |
Holboell's rock-cress, woodland rock-cress |
|
Habit | Perennials; long-lived; (often cespitose); sexual; caudex somewhat woody. | Perennials; short-lived; apomictic; caudex not woody. |
Stems | usually 2–5 per caudex branch, arising from margin of rosette near ground surface, (1–)1.5–4 dm, densely pubescent proximally, trichomes short-stalked, 5–8-rayed, 0.1–0.2 mm, moderately to sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 2–8(–9.6) dm, densely pubescent proximally, trichomes simple and short-stalked, 2–4-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1.5–3(–5) mm wide, margins entire, ciliate along petiole base, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 5–8-rayed 0.1–0.2 mm. |
blade oblanceolate, 2–8(–11) mm wide, margins entire or denticulate, ciliate proximally, trichomes (simple), 0.5–1 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed, 0.2–0.5 mm, these mixed with fewer simple ones. |
Cauline leaves | 5–14, not concealing stem; blade auricles 0.1–0.5 mm, surfaces of distalmost leaves pubescent. |
7–33, often concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves sparsely pubescent or glabrous. |
Racemes | 10–30-flowered, usually unbranched. |
15–63-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals usually white, 5–7 × 1–2 mm, glabrous; pollen ellipsoid. |
divaricate to pendent at anthesis; sepals pubescent; petals white to lavender, 5–6 × 1–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | ascending to divaricate-ascending, straight, 5–10 mm, pubescent, trichomes branched. |
reflexed, abruptly recurved near base, otherwise straight, 4–12 mm, sparsely pubescent or glabrous, trichomes spreading, simple and 2-rayed. |
Fruits | ascending to divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 3–5.5 cm × 0.9–1.2 mm; valves glabrous; ovules 56–84 per ovary; style 0.1–0.5 mm. |
reflexed to closely pendent, not appressed to rachis, not secund, straight or curved, edges parallel, (4.5–)5.5–8.5 cm × 1.5–2 mm; valves glabrous; ovules 70–110 per ovary; style 0.05–0.4 mm. |
Seeds | uniseriate, 0.8–1 × 0.6–0.9 mm; wing continuous (rarely absent), to 0.1 mm wide. |
uniseriate, 1.5–1.8 × 1.2–1.5 mm; wing continuous, 0.15–0.3 mm wide. |
2n | = 14. |
|
Boechera crandallii |
Boechera pinetorum |
|
Phenology | Flowering May–Jun. | Flowering May–Jul. |
Habitat | Rocky slopes and gravelly soil in sagebrush, mountain shrub, open conifer forests | Rock outcrops and gravelly soil in meadows and open conifer forests |
Elevation | 2000-2700 m (6600-8900 ft) | 1100-3200 m (3600-10500 ft) |
Distribution |
CO
|
CA; NV; OR
|
Discussion | Of conservation concern. Boechera crandallii is a sexual diploid known only from the Gunnison Basin of west-central Colorado. Despite its narrow range, the species appears to hybridize frequently, most notably with B. pallidifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Most authors (e.g., R. C. Rollins 1993; R. D. Dorn 2001; S. L. Welsh et al. 2003; N. H. Holmgren 2005b) have treated Boechera pinetorum as a variety of Arabis (Boechera) holboellii. Under this guise, the name has been applied to a vast array of plants collected throughout western North America. This includes a diversity of sexual diploids plus nearly every hybrid containing a genome from B. retrofracta. Based on re-examination of the type collection, we have adopted a much narrower concept of the species. Morphological evidence suggests that B. pinetorum is an apomictic triploid hybrid containing three different genomes, derived from B. rectissima, B. retrofracta, and B. sparsiflora. Plants closely resembling the type of B. pinetorum are confined to the northern Sierra Nevada and adjacent southern Cascades. The majority of collections previously associated with the epithet pinetorum represent B. pauciflora (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 372. | FNA vol. 7, p. 397. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis crandallii, Arabis stenoloba | Arabis pinetorum, Arabis divaricarpa var. pinetorum, Arabis holboellii var. pinetorum, B. holboellii var. pinetorum |
Name authority | (B. L. Robinson) W. A. Weber: Phytologia 51: 369. (1982) | (Tidestrom) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 271. (2007) |
Web links |
|