Boechera collinsii |
Boechera xylopoda |
|
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collin's suncress, Collins' rockcress |
beautiful rockcress, bigfoot hybrid rockcress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody. | Perennials; long-lived; apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
usually 1 from per caudex branch, arising from center of leaf tuft, elevated above ground surface on woody base, (3–)5–7.5 dm, pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.25–0.9 mm., similarly pubescent distally. |
Basal leaves | blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm. |
blade linear-oblanceolate to oblanceolate, 3–7 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 1.5 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.15–0.5 mm. |
Cauline leaves | (8–)15–40, concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent. |
8–18, usually not concealing stem; blade auricles 1–2 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | (15–)30–90-flowered, usually unbranched. |
20–38-flowered, usually unbranched. |
Flowers | descending to pendent at anthesis; sepals pubescent; petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous; pollen ellipsoid. |
divaricate to pendent at anthesis; sepals pubescent; petals purple, 9–12 × 2–3.5 mm, sparsely pubescent (often some trichomes abaxially); pollen spheroid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple. |
descending-divaricate, usually recurved, 9–20 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched. |
Fruits | strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm; valves glabrous; ovules 70–126 per ovary; style 0.2–0.5 mm. |
reflexed, not appressed to rachis, not secund, straight, edges parallel, 5–7 cm × 2–2.5 mm; valves glabrous proximally, sparsely pubescent distally; ovules 98–126 per ovary; style 0.1–0.5 mm. |
Seeds | uniseriate, 1–1.4 × 0.8–1 mm; wing continuous (rarely absent), to 0.1 mm wide. |
sub-biseriate, 2–2.5 × 1.3–1.6 mm; wing continuous, 0.2–0.25 mm wide. |
2n | = 14. |
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Boechera collinsii |
Boechera xylopoda |
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Phenology | Flowering May–Jun. | Flowering Mar–May. |
Habitat | Rocky and gravelly hillsides, prairies, open woods, floodplains | Rock outcrops and gravelly slopes, often under shelter of shrubs in desert scrub, sagebrush, and chaparral communities |
Elevation | 600-1400 m (2000-4600 ft) | 900-2000 m (3000-6600 ft) |
Distribution |
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
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CA; NV
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Discussion | Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although usually treated as a variety of Arabis (Boechera) pulchra, B. xylopoda is an apomictic taxon that is morphologically intermediate between B. perennans and B. pulchra (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison) and probably arose through hybridization between those two sexual diploids. Arabis trichopoda Greene (1908), not Turczaninow (1840) is an illegitimate name, sometimes found in synonymy with Boechera xylopoda. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 370. | FNA vol. 7, p. 411. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii | Arabis pulchra var. gracilis, Arabis pulchra var. glabrescens, Arabis pulchra var. viridis, B. pulchra var. gracilis |
Name authority | (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) | Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 250. (2007) |
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